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Carabidae Carabidae of the World

Subfamily Carabinae Latreille, 1802

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Endofallus Carabus (Morphocarabus) karpinskii Kryzhanovskij, Matveev, 1993Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
Carabus (Morphocarabus) illigeri illigeri Dejean, 1826 (as bjelasnicensis Apfelbeck, 1902)
Endophallus
endophallus as bjelasnicensis Apfelbeck, 1902
Copyright © Ruslan Panin
Carabus (Archicarabus) monticola fontanae Born, 1906Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
W Ukraine, Vinogradov distr.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
endophallus hampei
Vinogradov distr.
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) kollari Palliardi, 1825Carabus (Trachycarabus) estreicheri Fischer, 1822
Carabus (Trachycarabus) estreicheri Fischer, 1822
Endophallus
Lviv Region, Brody distr., near Yasenov vill.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus tscherkessicus Korge, 1964Carabus (Megodontus) imperialis Fischer, 1823
Carabus (Megodontus) imperialis Fischer, 1823
Endophallus
endophallus
Copyright © Safronov
Carabus (Megodontus) schoenherri sajanus Breuning, 1927Carabus (Tribax) titan titan Zolotarev, 1913Carabus (Tribax) steveni satanas Semenov, 1896 Carabus (Tribax) circassicus kubaniensis Semenov, 1896Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886Carabus (Trachycarabus) sibiricus rybinskii Reitter, 1886Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Carabus (Trachycarabus) sibiricus obliteratus Fischer, 1827
Endophallus
endophallus sibiricus obliteratus Fischer, 1827
NE Kazakhstan,
S Altai mts.,S Ulbinsky mt.h=500m.,Turgusun vill., 20.VI.2001
male 25mm.
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus fossulatus Dejean, 1826Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans
SW-Russia,
Rostov reg. env. stanitsa Razdorskaya, A.Ponomarev leg
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Carabus (Trachycarabus) sibiricus errans Fischer, 1823
Endophallus
endophallus sibiricus errans- Lugansk reg., near Slavyanoserbk
Copyright © Ruslan Panin
Carabus (Trachycarabus) scabriusculus scabriusculus Olivier, 1795Carabus (Trachycarabus) scabriusculus inapertus Motschulsky, 1850 (loc.typ)Carabus (Trachycarabus) perrini planus Gehin, 1885
Carabus (Trachycarabus) perrini planus Gehin, 1885
Endophallus
endophalus S Russia,Krasnodar reg., Anapa distr., B. Utrish vill., env.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) perrini planus Gehin, 1885
Carabus (Trachycarabus) perrini planus Gehin, 1885
Endophallus
endophalus Karacaj-Cherkessija, near Kart-Djurt vill., upper Kylan riv. h=1300m
Copyright © Ruslan Panin
Carabus (Trachycarabus) perrini perrini Dejean, 1831Carabus (Trachycarabus) mandibularis buchtarmensis Kryzhanovskij, 1953Carabus (Trachycarabus) besseri Fischer, 1822Carabus (Morphocarabus) wulffiusi wulffiusi Morawitz, 1862Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Carabus (Morphocarabus) illigeri simulator Kraatz, 1876
Endophallus
NE-Serbia, Krs Mts., Majdanpek env.,h=300m.20-31.V.2006, Pr.Sima leg
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus spasskianus Breuning, 1932Carabus (Morphocarabus) spasskianus schestopalovi Plutenko, 1995Carabus (Morphocarabus) scheidleri scheidleri Panzer, 1799Carabus (Morphocarabus) scheidleri ronayi Csiki, 1906 (? scheidleri)Carabus (Morphocarabus) scheidleri ronayi Csiki, 1906 (? scheidleri)Carabus (Morphocarabus) excellens Fabricius, 1798
Carabus (Morphocarabus) excellens Fabricius, 1798
Endophallus
endophallus (Morphocarabus) excellens excellens Moldova,
Anenii Noi distr., near Garbovets vill. 20.VI.08 Bodnar I.I.leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus)  helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male bronze colour.
Copyright © Ruslan Panin
Carabus (Morphocarabus)  helleri Ganglbauer, 1893
Carabus (Morphocarabus) helleri Ganglbauer, 1893
Endophallus
SE Moravia, Strani, V.2002, M.Snizek leg. male blue colour
Copyright © Ruslan Panin
Carabus (Tribax) apschuanus pseudoplatessa Gottwald, 1982Carabus (Archicarabus) montivagus montivagus Palliardi, 1825Carabus (Neoplectes) starckianus ganglbaueri Reitter, 1888Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844: 100Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Carabus (Morphocarabus) odoratus odoratus Motschulsky 1844 (as melleus Lapouge, 1909)
Endophallus
<p> endophallus odoratus odoratus (as melleus Lapouge, 1909)<br /> <br />     <br />  </p>
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus bargusinicus Shilenkov, 1996Carabus (Morphocarabus) monilis Fabricius, 1792
Carabus (Morphocarabus) monilis Fabricius, 1792
Endophallus
France 39, Foncina le Haut 300m
Copyright © Alexander Anichtchenko
Carabus (Morphocarabus) monilis Fabricius, 1792 (as interpositus Géhin, 1880)Carabus (Morphocarabus) mestscherjakovi shaman Shilenkov, 1996Carabus (Morphocarabus) kozhantschikovi kozhantschikovi Lutschnik, 1924Carabus (Morphocarabus) aeruginosus zyrjanovskianus Shilenkov & O.Berlov, 1996Carabus (Morphocarabus) aeruginosus aeruginosus Fischer, 1822Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Carabus (Eucarabus) ulrichi ulrichi Germar, 1824
Endophallus
endophallus ulrichi (as podolicus Semenov, 1898)
Copyright © Ruslan Panin
Carabus (Platycarabus) irregularis montandoni Buysson, 1882Carabus (Microplectes) riedeli Menetries, 1832Carabus (Microplectes) convallium convallium Starck, 1889Carabus (Microplectes) convallium argonautarum Semenov, 1898Carabus (Pachystus) tamsi tamsi Menetries, 1832Carabus (Pachystus) glabratus glabratus Paykull, 1790Carabus (Mesocarabus) cantabricus brabeus Schaufuss, 1876Carabus (Megodontus) violaceus fiorii Born, 1901Carabus (Megodontus) violaceus baeterrensis Lapouge, 1901Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Carabus (Megodontus) violaceus andrzejuscii Fischer, 1823
Endophallus
endophallus
Lviv reg., Nikolaev distr., near Krasiv vill.
Copyright © Ruslan Panin
Carabus (Megodontus) exaratus Quensel, 1806Carabus (Archicarabus) nemoralis nemoralis Muller, 1764Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Carabus (Platycarabus) fabricii malachiticus Thomson, 1875
Endophallus
Endophallus fabricii malachiticus
Copyright © Ruslan Panin
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Carabus (Neoplectes) daphnis daphnis Kurnakov, 1962
Endophallus
Georgia, Mingrelia, Lecheha vallis, Tehuri superior
Caucasus centr., 1200m, 27.6-7.7.1989 Retezar I. leg.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
Georgia, env. Martvilli, Askhi plateau
10-15.7.2001 A.Rubenian leg.
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Carabus (Neoplectes) daphnis askhicus Belousov, 1992
Endophallus
W Georgia, valley of Chkhorodukhu riv., 1000-1200m
14.V-6.VII.1991
Copyright © Alexander Anichtchenko
Carabus (Neoplectes) daphnis askhicus Belousov, 1992Carabus (Neoplectes) juenthneri avadcharensis Kurnakov, 1970Carabus (Neoplectes) reitteri laevisternis Starck, 1890Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
Carabus (Neoplectes) prometheus prometheus Reitter, 1887 (as nubicola Zolotarev, 1913)
Endophallus
Endophallus prometheus prometheus (as nubicola Zolotarev, 1913)
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Carabus (Neoplectes) prometheus prometheus Reitter, 1887
Endophallus
Endophallus prometheus prometheus Reitter, 1887
Copyright © Ruslan Panin
Carabus (Neoplectes) prometheus prometheus Reitter, 1887Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Carabus (Neoplectes) prometheus miroshnikovi Zamotajlov, 1990
Endophallus
aedeagus miroshnikovi Psebai env., Nikitino
Copyright © Ruslan Panin
Carabus (Neoplectes) kratkyi kratkyi Ganglbauer, 1890Carabus (Imaibius) nouristani nouristani (Ledoux, 1977)Carabus (Hygrocarabus) variolosus nodulosus Creutzer, 1799Carabus (Hygrocarabus) variolosus variolosus Fabricius, 1787Carabus (Eucarabus) manifestus pyonganicola Deuve & Li, 1998Carabus (Morphocarabus) praecellens x illigeri Dejean, 1826
Carabus (Morphocarabus) praecellens x illigeri Dejean, 1826
Endophallus
W-Croatia, Karlovac, near Bosiljevo vill., h~300m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Carabus (Morphocarabus) spasskianus aeruginosiformis Breuning, 1932
Endophallus
Russia, Khakassia,
W-Sayan, Alan Mt range, valley Enisey riv., 5 km S Mayna vill.,
Copyright © Ruslan Panin
Carabus (Trachycarabus) sibiricus gvozdevae Obydov, 1997Carabus (Morphocarabus) mestscherjakovi mestscherjakovi Lutshnik, 1924Carabus (Morphocarabus) spasskianus shoriensis Obydov, 1999Carabus (Neoplectes) starckianus obtusus Ganglbauer, 1886Carabus (Neoplectes) starckianus babukensis Zamotajlov, 1988 (as shchessiensis)Carabus (Carabus) granulatus granulatus Linne, 1758Carabus (Chrysocarabus) auronitens escheri Palliardi, 1825Carabus (Chrysocarabus) auronitens festivus Dejean, 1826Carabus (Chrysocarabus) auronitens farinesi Dejean, 1826Carabus (Chrysocarabus) hispanus Fabricius, 1787Carabus (Eucarabus) obsoletus obsoletus Sturm, 1815Carabus (Megodontus) violaceus aurolimbatus Dejean, 1829Carabus (Morphocarabus) spasskianus zinaidae Obydov, 1997Carabus (Morphocarabus) rothi hampei Kuster, 1846 (transition to rothi incompsus Kraatz, 1880)Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as  pseudopreyssleri Breuning, 1932)
Carabus (Morphocarabus) helleri Ganglbauer, 1893 (as pseudopreyssleri Breuning, 1932)
Endophallus
Czech Rep., NE Moravia, Moravian-Silesian Beskids Mt Rng, PR Trojacka, Hodslavice vill.,
Copyright © Ruslan Panin
Carabus (Morphocarabus) kollari Palliardi, 1825 (as semetricus Kraatz, 1878)Carabus (Morphocarabus) spasskianus geblerianus Obydov, 1999Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Carabus (Morphocarabus) rothi hampei (as ormayi Reitter, 1896) loc.typ.
Endophallus
near Beregovo, 27.IV-22.V..2011,, Panin R. & Chyzhevsky O. leg
Copyright © Ruslan Panin
Carabus (Morphocarabus) scheidleri ronayi Csiki, 1906 (? scheidleri)Carabus (Morphocarabus) praecellens sp.
Carabus (Morphocarabus) praecellens sp.
Endophallus
NC Croatia, Ivanščica Mts , Ivanec,4-18.VI.2011, Krause Rene leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Carabus (Morphocarabus) rothi hampei Kuster, 1846
Endophallus
C Romania, Tвrgu Mureş,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as zilahiensis Csiki, 1906)Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Carabus (Tomocarabus) bessarabicus ? concretus Fischer, 1823
Endophallus
Steppe Reserve “Provalsky",
Copyright © Ruslan Panin
Carabus (Neoplectes) satyrus duripshensis Kurnakov, 1972Carabus (Morphocarabus) regalis tschusovae Obydov, 2004Carabus (Semnocarabus) carbonicolor carbonicolor Morawitz, 1886Carabus (Sphodristocarabus) janthinus janthinus s.str. (as dvorshaki Mandl, 1975)Carabus (Neoplectes) apollo tenebricosus Kurnakov, 1962Carabus (Morphocarabus) rothi rothi Dejean, 1829
Carabus (Morphocarabus) rothi rothi Dejean, 1829
Endophallus
C Romania, Sibiu Area, right bank Olt River, Bradu vill., h=380 m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) venustus venustus Morawitz, 1862Carabus (Pachystus) hungaricus cribellatus Adams, 1812Carabus (Pachystus) hungaricus scythus Motschulsky, 1847Carabus (Trachycarabus) sibiricus sibiricus Fischer, 1822Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Carabus (Mesocarabus) dufouri dufouri Dejean, 1829
Endophallus
Jaen reg., Monte Javalcuz h=1436 m
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Carabus (Mesocarabus) lusitanicus brevis (as sorianus Jeanne, 1972)
Endophallus
NC-Spain, Soria, Puerto de Oncala
Copyright © Ruslan Panin
Carabus (Carabus) arvensis arvensis Herbst, 1784
Carabus (Carabus) arvensis arvensis Herbst, 1784
Endophallus
W Ukraine, NW Podolian, Lviv reg., Javoriv distr.,near Stradch vill.,
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Carabus (Tribax) puschkini ishikawaianus Breuning & Ruspoli, 1970
Endophallus
NE-Turkey: Artvin:Hopa, Borзka env., h=450 m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Carabus (Morphocarabus) praecellens praecellens Pillardi, 1825
Endophallus
Serbia, Fruska Gora Mts, 20-31.V.2006, M.Sima leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Carabus (Morphocarabus) illigeri versicolor Frivaldsky, 1835
Endophallus
C Bulgaria: Stara planina Mts, Gabrovo
Copyright © Ruslan Panin
Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Carabus (Tribax) puschkini ponticus Deyrolle, 1869
Endophallus
NE-Turkey:Trabzon, Zigana Pass h=1700m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Carabus (Morphocarabus) rothi comptus (as szoerenyensis Csiki, 1908)
Endophallus
SW Romania, Tarcu Mts, vf. Nedeia h=2000m,
Copyright © Ruslan Panin
Carabus (Megodontus) bonvouloiri bonvouloiri (as kaschkarensis Blumenthal & Breuning, 1967)Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Carabus (Morphocarabus) rothi hampei (as gutiiensis Takács & Lie, 1992)
Endophallus
N Romania, E Carpathian, Maramureş Reg., Gutai Mts, cabana Mogosa, 4 km N Baia Sprie,
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Carabus (Sphodristocarabus) macrogonus macrogonus Chaudoir, 1847
Endophallus
NE-Turkey:Trabzon env., Sumela,
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Carabus (Sphodristocarabus) macrogonus pseudokolatensis Heinz & Korge, 1967
Endophallus
NE-Turkey::Kackar Dagi Mt, Ayder S of Ardesen h=1600m
Copyright © Ruslan Panin
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Carabus (Leptocarabus) seishinensis seishinensis Lapouge, 1931
Endophallus
NE-China:Liaoning prov.,Fengcheng City, Miaoyang,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Carabus (Morphocarabus) rothi incompsus Kraatz, 1880
Endophallus
C Romania, Brașov County, Cristian vill., h=380 m
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Carabus (Morphocarabus) rothi alutensis Savulescu, 1972
Endophallus
C Romania, S Carpathians, Rimnicu Vilcea env., Calimaneşti, Cozia Mts.,
Copyright © Ruslan Panin
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Carabus (Lamprostus) robustus ducalis Schaufuss, 1882
Endophallus
NE-Turkey:Trabzon, Zigana Pass h=2000m
Copyright © Ruslan Panin
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Carabus (Diocarabus) fraterculus fraterculus Reitter, 1895
Endophallus
N-Korea, N Pyongan Prov.,Sakju, Mt Ponsan,
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) armeniacus Mannerheim, 1830
Carabus (Sphodristocarabus) armeniacus Mannerheim, 1830
Endophallus
NE Turkey, Artvin Prov., Şavşat, Зamlıca.
Copyright © Ruslan Panin
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Carabus (Pachystus) hungaricus hungaricus Fabricius, 1792
Endophallus
Czech, S Moravia, Pouzdrany
Copyright © Ruslan Panin
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Carabus (Lamprostus) torosus torosus Frivaldsky, 1835
Endophallus
NE-Bulgaria, Dobrich Region, Kavarna env.,
Copyright © Ruslan Panin
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Carabus (Semnocarabus) erosus kuldshaensis Ballion, 1878
Endophallus
S-Kazakhstan:Ili River valley,:near Bakanas,
Copyright © Ruslan Panin
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Carabus (Megodontus) stroganowi transfuga Semenov, 1898
Endophallus
SE-Azerbaijan: Talysh, Kalvaz,
Copyright © Ruslan Panin
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Carabus (Deroplectes) dokhtouroffi dokhtouroffi Ganglbauer, 1886
Endophallus
NC Tadzhikistan, Pamir, N-slope Peter I.Mt Rng, h=2100m, Ganishou vill.
Copyright © Ruslan Panin
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017
Carabus (Morphocarabus) viridilimbatus panini I.Solodovnikov 2017 Holotype
Endophallus
Russia, SE Yakutia, Neryungri distr., Tokinski Stanovik Mt Rng., Beder riv., (right tributary rivers Algama), 26.VI.1996
Copyright © Ruslan Panin
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Carabus (Ulocarabus) stschurovskii Solsky, 1874
Endophallus
Tadzhikistan, Pamir-Alay, S slope Gissar Mt.Range, Takob, h-1700m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Carabus (Morphocarabus) rothi hampei (as diffinis Csiki, 1906)
Endophallus
NW Romania, Apuseni Mts, Trascău Mt Rng, Petresti de Jos, 12 km E Turda,.
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus Reitter, 1895
Carabus (Ulocarabus) theanus Reitter, 1895
Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
Copyright © Ruslan Panin
Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Carabus (Mesocarabus) (? lusitanicus) gougeleti Reiche, 186
Endophallus
CW Spain, Province Huelva, Arroyomolinos de Leon.
Copyright © Ruslan Panin
Carabus (Platycarabus) fabricii ucrainicus Lazorko, 1951Carabus (?Oreocarabus) titanus Breuning, 1933
Carabus (?Oreocarabus) titanus Breuning, 1933
Endophallus
C-China Sichuan prov., Nanping 1500m,
Copyright © Ruslan Panin
Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Carabus (Megodontus) kolbei kolbei Roeschke, 1887
Endophallus
N-Japan, C-Hokkaido, Hidaka distr., Niikappu env.,
Copyright © Ruslan Panin
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Carabus (Leptocarabus) procerulus Chaudoir, 1862
Endophallus
Japan:S Honshu , Matsubara-ko,
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Carabus (Trachycarabus) latreillei ?semicoriaceus Kraatz, 1881
Endophallus
NE-China: Liaodong Peninsula, Liaoning prov., Gaizhou distr., Mt. Buyun Shan.
Copyright © Ruslan Panin
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Carabus (Carabus) arvensis hokkaidoensis Lapouge, 1924
Endophallus
N Japan:Hokkaido, Hamamasu env.,
Copyright © Ruslan Panin
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Carabus (Ohomopterus) japonicus japonicus Motschulsky, 1857
Endophallus
S Japan, Kyushu, Kumamoto Pref., Izumi viil., h=600 m
Copyright © Ruslan Panin
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Carabus (Eurycarabus) famini fezzanus Lapouge, 1915
Endophallus
Morocco, Middle Atlas Mt Rng, Azrou city h=1500m,
Copyright © Ruslan Panin
Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Carabus (Leptocarabus) kumagai kumagai Kimura & Komiya, 1974
Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
Copyright © Ruslan Panin
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Carabus (Oreocarabus) taedatus agassi Le Conte, 1850
Endophallus
SC-USA, New Mexico, Santa Fe,
Copyright © Ruslan Panin
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Carabus (Ohomopterus) yamato kitai (Imura, 2004)
Endophallus
Japan, S Honshu, Mie Pref., Hakusan,
Copyright © Ruslan Panin
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Carabus (Trachycarabus) latreillei latreillei Fischer, 1822
Endophallus
N Mongolia, Khövsgöl reg., Lake Khovsgol, h=1640m, 51*:12’N; 100*:45’E, steppificated sites,
Copyright © Ruslan Panin
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Carabus (Ulocarabus) theanus theanus Reitter, 1895
Endophallus
Tadzhikistan, S slope Gissar Mt.Range, left bank of the river Sorbo, Haidar gorge,
Copyright © Ruslan Panin
Carabus (Eucarabus) manfredeggeri Müller 2012Carabus (Imaibius) dardiellus perobscurus Deuve, 1980Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Carabus (Morphocarabus) illigeri illigeri (as tarensis Toševski, 1990)
Endophallus
WC-Serbia: National park Tara Mts.:NE Mitrovac h=1500m,
Copyright © Ruslan Panin
Carabus (Cupreocarabus) huangi huangi Deuve, 1992Carabus (Megodontus) violaceus dryas Gistl, 1857
Carabus (Megodontus) violaceus dryas Gistl, 1857
Endophallus
NE-Greece, Mt. Olympos, Litochoro env.
Copyright © Ruslan Panin
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Carabus (Lamprostus) saulcyi saulcyi Piochard, 1875
Endophallus
C Lebanon, Jbeil Distr., Ehmej vill., h=1100 m
Copyright © Ruslan Panin
Carabus (Pachystus) hortensis hortensis Linne, 1758
Carabus (Pachystus) hortensis hortensis Linne, 1758
Endophallus
W-Ukraine, Male Polissya, near Kamenka-Buzka vill.
Copyright © Ruslan Panin
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Carabus (Oreocarabus) guadarramus La Ferte-Senectere, 1847
Endophallus
WC Spain, Cáceres, Hervás, h=1000m
Copyright © Ruslan Panin
Carabus (?Oreocarabus) qinlingensis Imura, 1993
Carabus (?Oreocarabus) qinlingensis Imura, 1993
Endophallus
China, W-Shaanxi, Taibai Shan Mts, btw.Houzhenzi/Ba Xian Tai, nr.Nan Tian Men h=3200m, 33°55'17"N/107°46'53"E,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Carabus (Morphocarabus) rothi hampei (as hunyadensis Csiki, 1926)
Endophallus
CW Romania, Apuseni Mts, Metalliferous Mt Rng, Hunedoara reg., near Săcărâmb vill.
Copyright © Ruslan Panin
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Carabus (Oreocarabus) latro minshanensis Deuve, 1987
Endophallus
China, N-Sichuan, Jiuzhaigou env.,
Copyright © Ruslan Panin
Carabus (Meganebrius) quinlani smetanai Häckel, 2011
Carabus (Meganebrius) quinlani smetanai Häckel, 2011 Paratype
Endophallus
C-Nepal:Dhaulagiri Massif:Baglung Distr.,(1.5km W of Okhale vill., 28.32’152’’N/ 83.48’673’E, h=2600m)
Copyright © Ruslan Panin
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Carabus (Rhigocarabus) ladygini degensis Deuve, 1989
Endophallus
C-China, NW- Sichuan prov., Garzê Tibetan AP, Pass Garzê – Sêrtar
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus azurescens Dejean, 1826
Carabus (Megodontus) violaceus azurescens Dejean, 1826
Endophallus
W-Croatia, Velebit Mts., betw.Karlobag & Brusane vill.,
Copyright © Ruslan Panin
Carabus (Megodontus) violaceus germarii Sturm, 1815
Carabus (Megodontus) violaceus germarii Sturm, 1815
Endophallus
N Italy, Trent, Ampola, Mt. Tremalzo h=1700m
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Carabus (Morphocarabus) zawadzkii seriatissimus Reitter, 1896
Endophallus
W Ukraine,Transcarpathian Reg., Rakhiv distr., Marmarosh Mts., near Lug vill., h=400 m.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
Transcarpathian region, Uzhhorod distr., 28.IV-23.V.2011
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
SE Slovakia, Slanske vrchy Mts, Herlianske sedlo, 8-19.VI.2009, Krause Rene leg.
det.Panin R.
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
Gorgany Mts. h=700 m, 16.VI.2011,
Copyright © Ruslan Panin
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Carabus (Morphocarabus) zawadzkii zawadzkii, Kraatz 1854
Endophallus
W.UKRAINE, Ukrainian Carpathians
Lviv Reg., Skole distr., Skolivski Beskidy Mts.
Copyright © Igor Solodovnikov
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
Carabus (Mesocarabus) cantabricus cantabricus Chevrolat, 1840
Endophallus
NW-Spain, Pontevedra prov., A Cañiza viil., Alto de Fontefría h=800m
det.Panin R.
Copyright © Ruslan Panin
Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Carabus (Mesocarabus) dufouri baguenai Breuning, 1926
Endophallus
SE Spain, Almería Province, Carboneras env.
det. R.Panin
Copyright © Ruslan Panin
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Carabus (Mesocarabus) macrocephalus Dejean, 1826
Endophallus
NW-Spain,Cantabrian Mts, Picos de Europa Mt Rng, Puerto de Panderruedas h=1460 m,
Copyright © Ruslan Panin
Carabus (Carabus) arvensis deyrollei Gory, 1839
Carabus (Carabus) arvensis deyrollei Gory, 1839
Endophallus
NW-Spain, Cantabria, Udías env., Monte Corona h=100m
Copyright © Ruslan Panin
Carabus (Mesocarabus) latus latus Dejean, 1826
Carabus (Mesocarabus) latus latus Dejean, 1826
Endophallus
C-Spain, Toledo, Mts de Toledo, Sierra del Castañar Rng, Puerto del Milagro env, h=920m
Copyright © Ruslan Panin
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Carabus (Semnocarabus) carbonicolor korolkowi Semenov, 1896
Endophallus
NC-Kyrgyzstan, Kyrgyz Mt.Rng , 10 km SW of Bishkek, Chon-Aryk env., h=1100m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906
Endophallus
N Hungary, Pilis Mts., Piliszentlelek,20-27.VII.2011, Krause Rene leg.
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as pseudojucundus Retezár, 1974)
Endophallus
NW-Hungary::Komarom distr., Tardos, Gerecse Mts.
Copyright © Ruslan Panin
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Carabus (Morphocarabus) praecellens jucundus Csiki, 1906 (as vertesensis Retezár, 1974)
Endophallus
NW Hungary: Fejer: Vertes Mt Rng, Tatabanya
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Carabus (Mesocarabus) lusitanicus helluo (as albarracinus Ganglbauer, 1886)
Endophallus
CE-Spain, Provincia Teruel, Sierra de Javalón Rng, El Cañigral env., h=1460m,
Copyright © Ruslan Panin
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Carabus (Mesocarabus) lusitanicus castilianus Dejean, 1826
Endophallus
CW-Spain:Salamanca, Brincones env., h=750m
Copyright © Ruslan Panin
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Carabus (Mesocarabus) problematicus harcyniae (as belgicus Lapouge, 1913)
Endophallus
Luxembourg, Bollendorf-Pont env., 3.III.1999
Copyright © Ruslan Panin
Carabus (Mesocarabus)  (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Carabus (Mesocarabus) (? problematicus trans lusitanicus) planiusculus (as xaxarsi Born, 1917)
Endophallus
NE Spain, Catalonia,Girona Prov., San Hilario Sacalm env.<br />
Copyright © Ruslan Panin
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Carabus (Aulonocarabus) kurilensis ? daisetsuzanus Kono, 1936
Endophallus
N-Japan, Hokkaido, Katō District, Kamishihoro, Horoka near Maruyama Bridge,
Copyright © Ruslan Panin
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Carabus (Tribax) agnatus pseudagnatus Novotný & Voříšek, 1988
Endophallus
W-Caucasus: Abkhasia, Gudauta distr., Mt Anchkho h=2200-2350 m,
Copyright © Ruslan Panin
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Carabus (Neoplectes) edithae exedithae Gottwald, 1985
Endophallus
NС-Caucasus, Karachaevo-Cherkessia, right bank Pkhiya river, Abishir-Akhuba Mt.Range h=2250-2600 m,
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Carabus (Sphodristocarabus) separatus persa Lapouge, 1924
Endophallus
N-Iran: Mazandaran Province, Elborz Mt.Rng, Vandarbon env.
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Carabus (Morphocarabus) rothi comptus (as borlovensis Lie, 1994)
Endophallus
SW Romania,, Banat, Borlova env.,
Copyright © Ruslan Panin
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Carabus (Limnocarabus) clathratus auraniensis Muller, 1902
Endophallus
W Ukraine, Transcarpathian Reg., 4 km NE Chop, forest Peresh.
Copyright © Ruslan Panin
Carabus (Morphocarabus) brinevi Kabak, 2014Carabus (Ophiocarabus) nilkiensis Kabak, 2014Carabus (Mimocarabus) pumilio Kuster, 1846
Carabus (Mimocarabus) pumilio Kuster, 1846
Endophallus
S-Armenia, Vayots Dzor Region, E Ughedzor vill., h=2630 m
Copyright © Ruslan Panin
Carabus (Pachycarabus) staehlini Adams, 1817
Carabus (Pachycarabus) staehlini Adams, 1817
Endophallus
S Russia, E Caucasus, Dagestan, Nukatl Mt Rng h=2600m,
Copyright © Ruslan Panin
Carabus (Imaibius) boysi boysi Tatum, 1851
Carabus (Imaibius) boysi boysi Tatum, 1851
Endophallus
NW-India, Himachal Pradesh, Manikaran env., Pulga vill.
Copyright © Ruslan Panin
Carabus (Carabus) shamaevi longchi Kleinfeld & Imura, 2009Carabus (Neoplectes) besleticus resheviensis Solodovnikov, Zamotajlov & Fominykh, 2014Carabus (Neoplectes) satyrus Kurnakov, 1962Carabus (Neoplectes) pseudopshuensis Zamotajlov, 1991Carabus (Neoplectes) besleticus besleticus Kurnakov, 1972Carabus (Neoplectes) besleticus mtsaranus Kurnakov, 1972Carabus (Neoplectes) besleticus dsychvensis Gottwald, 1985Carabus (Neoplectes) besleticus duripshensis Kurnakov, 1972Carabus (Neoplectes) besleticus napraensis Belousov & Zamotajlov, 1993Carabus (Neoplectes) besleticus anastasiae Solodovnikov, 2014Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Carabus (Lamprostus) torosus sinopensis Breuning & Ruspoli, 1970
Endophallus
NC-Turkey, Sinop province, Erfelek env., h=200m
Copyright © Ruslan Panin
Carabus (Archicarabus) phoenix Lapouge, 1924
Carabus (Archicarabus) phoenix Lapouge, 1924
Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
Copyright © Ruslan Panin
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Carabus (Procrustes) piochardi praestigiator Morawitz, 1886
Endophallus
C-Lebanon, Byblos /Jbeil Distr., Kfardebian env., h=1200m,
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) blumenthaliensis aequicostalis Staven & Heinz, 1995Carabus (Mesocarabus) riffensis Fairmaire, 1875
Carabus (Mesocarabus) riffensis Fairmaire, 1875
Endophallus
Morocco, Rif, Foresta di Ketama, 25.06.1998, 1600m, leg. D. Sechi (immature)
Copyright © Alexander Anichtchenko
Carabus (Ophiocarabus) wusunshanicus Kabak, 2015Carabus (Ophiocarabus) regeli regeli Morawitz, 1886Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Carabus (Lamprostus) hemprichi elonensis Schweiger, 1970
Endophallus
N-Israel, Midlle Galilea, Nazareth env., Hoshaya viil.
Copyright © Ruslan Panin
Carabus (Procrustes) paiafa paiafa White, 1845Carabus (Carabus) stscheglowi Mannerheim, 1827
Carabus (Carabus) stscheglowi Mannerheim, 1827
Endophallus
Russian, Chuvashia, Cheboksary, Zavolzhye distr., , near lake Astrakhanka,
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Carabus (Morphocarabus) odoratus baeri Menetries, 1851: 45
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Carabus (Morphocarabus) zawadzkii dissimilis Csiki, 1906
Endophallus
NE-Hungary, 4 km S Sátoraljaújhely, right bank riv. Bodrog (Long-erdő),
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Carabus (Morphocarabus) rothi distinguendus Csiki, 1906
Endophallus
NC-Hungary: E- part of Budapest,Cinkota,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Carabus (Morphocarabus) rothi distinguendus (as bieneri Mandl, 1965)
Endophallus
C-Slovakia, Detva Distr., Stara Huta env. h=700-800m,
Copyright © Ruslan Panin
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Carabus (Morphocarabus) rothi distinguendus (as subparvulus Mandl, 1965)
Endophallus
NC-Hungary:Borzsony Mts, Nograd-Diosjeno,
Copyright © Ruslan Panin
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966
Carabus (Lamprostus) erenleriensis erenleriensis Schweiger, 1966
Endophallus
NW-Turkey:Bolu Prov., Abant Lake
Copyright © Ruslan Panin
Carabus (Sphodristocarabus) adamsi Adams, 1817
Carabus (Sphodristocarabus) adamsi Adams, 1817
Endophallus
CN-Georgia, N of Tbilisi, NW of Tianeti, Tvalivi env.,
Copyright © Ruslan Panin
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017
Carabus (Morphocarabus) odoratus kurmaevi I.Solodovnikov, 2017 Holotype
Endophallus
Russia, Siberia, Krasnoyarsk Krai, the North Yenisei district, 55 km of CE of the vill. Bryanka, valley of the river Chirimba, stream Nakhodny, h = 350 m, the mixed forest (birch and pine); on a steep slope along an animal track on a hill brow, 23.VII-20.VIII.2015, leg. A.V. Kurmaev
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017
Carabus (Morphocarabus) viridilimbatus saluki I.Solodovnikov 2017 Paratype
Endophallus
Russia, SW Taymyr, near Talnakh, 25 km N Norilsk, hilly tundra, hillside overgrown with alder slate, dwarfish birches, blueberry, h = 150-250 м, 24-28.06.2003, leg. S.Saluk
Copyright © Igor Solodovnikov
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Carabus (Morphocarabus) viridilimbatus viridilimbatus Motschulsky 1859
Endophallus
Russia, Chita Region, Udokansky district. Kodar Range, 56 °34'42,9 " N, 118 °29'34,95" E, 6/13/2002, leg. A. Pokhorov
Copyright © Igor Solodovnikov
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