Nebria uralensis Glasunov, 1901 Male adult
Bashcortostan., mountain Bolshoy Iremal, alt. 1580 m. 19/08/2008.
Leg. E.Chibilov
Copyright ©
Sunvas 
Leistus terminatus Panzer, 1793 Male adult
Copyright ©
Ruslan Panin 
Nebria (Nebriola) gosteliae Huber, 2010 Paratype
Male adult
Paratype: Lago Mucrone, Oropa, Biella, 27. 6. 2006, leg. A. Szallies. Body size: 9 mm. Illustration: Hans-Peter Wymann (Jegenstorf).
Copyright © - - -
Nebria (Nebria) tenella tenella Motschulsky, 1850 Female adult
NW-Caucasus: Karachay-Cherkessia, Abishira–Akhuba Mnt. Range h=2000-2650m, Kyafar-Agur Riv. 14–17.VII.2009, Fominykh D. leg.
Copyright ©
Ruslan Panin 
Leistus (Leistus) baenningeri Roubal, 1926 Male adult
East Carpathians, Ukraine Marmarosh Mts. h=1500 16.VII.1987 Leg/Det. Rizun V.
male, 8 mm.
Copyright ©
Ruslan Panin 
Leistus (Leistus) baenningeri Roubal, 1926 Male adult
W UKRAINE, NE Carpathians Mts. Transcarpathian reg., Rakhov distr., Chornagora Mt range, N-slope of Sheshul Mt. h=1450m. 9.VI.2009 Panin R.leg./
L=7mm
Copyright ©
Ruslan Panin 
Leistus (Leistus) ferrugineus Linne, 1758 Male adult
Moskow Reg., Istra Distr., E shore of Istrinskoe Reserv., Novorakovo Vill. 16.IX.2007 K.Makarov leg.
L=6.8 mm
Copyright ©
Kirill Makarov 
Leistus (Leistus) piceus piceus Frolich, 1799 Male adult
W.Ukraine,
Transcarphatian reg., Rakhov distr., Chornogora Mt range h=1850m. 28.VI-9.VIII.2008, Panin R.leg.
male, 8.5mm.
Copyright ©
Ruslan Panin 
Leistus (Leistus) piceus piceus Frolich, 1799 Female adult
NW Ukrainian Carpathians
Lviv reg., Staryi Sambor Distr., near Dobromyl , h=500m 6-28.VI.2009, Panin R. leg.
photo Panin R.
Copyright ©
Ruslan Panin 
Leistus (Leistus) terminatus Panzer, 1793 Female adult
length: 7,7 mm
West Siberia, KHMAO-Yugra, Khanty-Mansiysk distr., Ob river, Bolshoy Tchukhtinsiy island
4.08.2007.
Zinoviev E.leg
Copyright ©
Evgeniy Zinoviev 
Leistus (Pogonophorus) rufomarginatus Duftschnnd, 1812 Male adult
W. UKRAINE, NW Podolian
Ternopil Reg., Pidvolochisk distr., Forest Bogdanivskyi, 2.VI-15.VII.2010 Panin R. leg.
9.5 mm
Copyright ©
Ruslan Panin 
Nebria (Alpaeonebria) dilatata Dejean, 1831 Male adult
Tenerife, Mte Esperanza, 7.11.1980 J. de Ferrer leg.
14 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Alpaeonebria) fuscipes Fuss, 1850 Male adult
Ukrainian Carpathians
Transcarpathian reg., Rakhov Distr., Chornogora Mts., h~1650m., 29.VI-28.VII.2007, Panin R. leg.
male, 13 mm
Copyright ©
Ruslan Panin 
Nebria (Alpaeonebria) reitteri Rybinski, 1902 (loc.typ.) Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora Mts. h~1650 m, 29.VI-28.VII.2007, Panin R. leg.
L=12.5 mm
Copyright ©
Ruslan Panin 
Nebria (Boreonebria) gyllenhali castanipes Kirby, 1837 Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright ©
Henri Goulet 
Nebria (Boreonebria) gyllenhali gyllenhali Schonherr, 1806 Male adult
Ukrainian Carpathians
Zakarpatskaya Prov., Rakhov Distr., Chornogora Mts. Sheshul h=1450 m Panin R.leg. 28.VII.2007
Copyright ©
Ruslan Panin 
Nebria (Boreonebria) heegeri Dejean, 1826 Male adult
W. Ukraine,
Ivano-Frankivsk reg., Yaremcha distr., Carpathians, Gorgany mts., h=700m. Panin R. leg/det. 23.VI.08
male 11 mm
Copyright ©
Ruslan Panin 
Nebria (Boreonebria) nivalis gaspesiana Kavanaugh Male adult
Nebria (Boreonebria) nivalis gaspesiana
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright ©
Henri Goulet 
Nebria (Boreonebria) subdilatata Motschulsky, 1844 Male adult
SE Siberia, Irkutsk Area, Khamar-Daban Mountains,6-8.VI.2007 D.Fominyh leg.
L=10mm.
Copyright ©
Ruslan Panin 
Nebria (Catonebria) aenea aenea Gebler, 1825 Male adult
Altai, Kuraiskii mt.rng., Kuraika riv.,
1800m 6.8.1994 R. Dudko leg.
12 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) catenulata Fischer von Waldheim, 1820 Male adult
E Sayan, Tunkinskie goltsy, up of Kyngarga riv.
1500m 18-23.7.1995 Anichtchenko A. leg.
12 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) fulgida Gebler, 1847 Female adult
Buryatia, Hamar-Daban mts., Osinovka Mishihinskaya riv.,
2000m 19.7.1995
Anichtchenko A. leg.
11 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) fulgida Gebler, 1847 Female adult
Hamar-Daban mts. Komarinskii mt. rng,
Podkomarnaya riv., 1600m, 10-14.6.1996
Anichtchenko A. leg.
11mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) gebleri Dejean, 1831 Female adult
USA Montana, Mineral Co. Trovd Creec
13.5 mil NE of Hoodoo pass 5.8.1996
11 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) kincaidi Schwarz, 1900 Male adult
USA Olympic Mts., Jefferson Co.
Washington 7.2005 Maynard leg.
11 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) piperi Van Dyke, 1925 Female adult
USA Pierce Co., Washington,
Tahoma riv., 07.2005 C.Mayard leg.
19 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) vandykei Banninger, 1928 Lectotype Male adult
Museum of Zurich
photo Anichtchenko
Copyright ©
Alexander Anichtchenko 
Nebria (Eonebria) djakonovi Semenov et Znojko, 1928 Male adult
Far East, Lazovskij Nat. Res., Formozova spring 43°00'06"N 134°05'13"E 10.VII.2005 K.Makarov leg.
L=11.0 mm
Copyright ©
Kirill Makarov 
Nebria (Eunebria) nigerrima Chaudoir, 1846 Male adult
Karachaevo-Cherkessia, Zelenchukskii distr., near Bukovo vill.,h=2000m., 9-13.VII.2007, S.Dementiev leg. det. Anichtchenko A. 2018
12mm.
Copyright ©
Ruslan Panin 
Nebria (Eunebria) jockischii hoepfneri Dejean, 1826 Male+Female adult
W. Ukraine, NE Carpathians,
Ivano-Frankivsk reg., Yaremcha distr., Gorgany mts., h=650m, Panin R. leg. 3.VIII.2008
female, 14 mm
Copyright ©
Ruslan Panin 
Nebria (Eunebria) lewisi Bates, 1874 Male adult
Japan, Ehime pref., Kamo-ike, Kikuma
Imabari 30.10.2005 T.Kurihara leg.
12 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Eunebria) picicornis radjabii Morvan, 1973 Paratype Female adult
Museum of Zurich
photo Anichtchenko
Copyright ©
Alexander Anichtchenko 
Nebria (Eunebria) yunnana Banninger, 1928 Holotype Male adult
Museum of Zurich
photo Anichtchenko
Copyright ©
Alexander Anichtchenko 
Nebria (Eurynebria) complanata Linne, 1767 Male adult
length: 17 mm<br />
France<br />
leg.det. O.Schmitt<br />
Foto: U.Schmidt, 2008
Copyright ©
U.Schmidt 
Nebria (Nakanebria) gibbulosa gibbulosa Motschulsky, 1860 Male adult
Kunashiri Isl., hills N of Severyanka River mouth 44°19'32"N 146°00'03 E h~200 26.VI.2008 leg.K.Makarov
L=11.0
Copyright ©
Kirill Makarov 
Nebria (Nebria) bonellii M.F. Adams, 1817 Male adult
NW Caucasus, Adygeya, Psheho-Su Mount. env. h=1500 m, 07.07.2006, D. Safronov leg.
Copyright ©
Safronov 
Nebria (Nebria) brevicollis Fabricius, 1792 Male adult
W.UKRAINE, Ternopol Reg., Pidvolochisk distr.,Supranivka vill., Panin R.Leg. 14.VI.08
long =12 mm
Copyright ©
Ruslan Panin 
Nebria (Nebria) commixta Chaudoir, 1850 Male adult
NW-Caucasus, Krasnodar reg., Mostovskoj distr., Skirda Mts. Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008 D. Fominyh leg.
long - 14mm.
Copyright ©
Ruslan Panin 
Nebria (Nebria) schlegelmilchi M.F. Adams, 1817 Female adult
Armenia,
Stepanavan, Pushkin Mountain Pass h=2000m, 18-19.IX.1989,Kazaryan P.V. leg.
14mm
Copyright ©
Ruslan Panin 
Nebria (Nebria) transsylvanica Germar, 1824 Female adult
Ukrainian Carpathians,
Transcarpathian reg., Rakhov Distr., Chornogora Mts. h~2000m., Panin R. leg.
L=12mm
Copyright ©
Ruslan Panin 
Nebria (Nebria) retrospinosa Heyden, 1885 Female adult
NW-Caucasus, Krasnodar reg., Mostovskoj distr., Skirda Mt.Range, Yatyrgvart Mt., h=2300-2500m. 09-10.VII.2008 D. Fominyh leg.
L=13mm.
Copyright ©
Ruslan Panin 
Nebria (Nebria) viridipennis Reitter, 1885 Male adult
Karachaevo-Cherkesia, near Alibek
10-11.6.1996 Arzanov Yu. leg.
13 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Nebriola) heeri praegensis Huber et Molenda, 2004 Holotype
Male adult
"Seehalde" bei Prдg (Gemeinde Todtnau, Deutschland), 600 m ь. M., 23.
6. 1997 (Fallenfang), leg. R. Molenda, deponiert am Naturhistorischen Museum Bern (NMBE).
male
as cordicollis praegensis Huber et Molenda, 2004
Copyright © - - -
Nebria (Paranebria) livida livida Linne, 1758 Male adult
Copyright ©
Ruslan Panin 
Nebria (Reductonebria) acuta Lindroth, 1961 Paratype Male adult
Museum of Zurich
photo Anichtchenko A.
Copyright ©
Alexander Anichtchenko 
Nebria (Reductonebria) appalachia Darlington, 1931 Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright ©
Igor Sokolov 
Nebria (Reductonebria) appalachia Darlington, 1931 Paratype Male adult
Museum of Zurich
photo Anichtchenko A.
Copyright ©
Alexander Anichtchenko 
Nebria (Reductonebria) pallipes Say, 1823 Male adult
Great Smoky Mountains National Park
photo I. M. Sokolov
Copyright ©
Igor Sokolov 
Nebria (Reductonebria) pallipes Say, 1823 Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright ©
Henri Goulet 
Nebria (Reductonebria) suturalis LeConte, 1850 Male adult
Canada
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright ©
Henri Goulet 
Nebria (Tyrrhenia) rubicunda rubicunda Quensel, 1806 Male adult
Spain, Andalucia, Sierra de Almijara
3 km O de Otivar, rio Verde, 13.3.2005
Anichtchenko A. leg.
11 mm
Copyright ©
Alexander Anichtchenko 
Nebria (Vancouveria) virescens Horn, 1870 Male adult
Canada,
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright ©
Henri Goulet 
Nebria (Nippononebria) changbaiensis Kavanaugh & Liang, 2010 Holotype Male adult
Jilin Province, Changbaishan, Tianchi, waterfall, 2000–2600m, Chinese Academy of Sciences; 1987.VII.22, Guiyun Deng collector
male
Scale line = 1.0 mm.
deposited in IOZ
Copyright ©
Alexander Anichtchenko 
Nebria (Catonebria) banksi Crotch, 1871 Male adult
Copyright ©
Kirill Makarov 
Nebria (Boreonebria) heegeri Dejean, 1826 Female adult
W-UKRAINE, Iv. Frankivsk reg.,
Yaremche distr., Gorgany Mts, on the bank of Prutets river h =700м., 24.VI.2011,
Panin R. leg.
L=11 mm
Copyright ©
Ruslan Panin 
Nebria (Catonebria) baicalopacifica Dudko et Shilenkov, 2006 Female adult
Jewish Aut. Reg., Obluchie Distr., 3-5 km upper Radde Vill. on Lagar Riv., valley 21.07.2004 I.Melnik leg.
L=12.0 mm
Copyright ©
Kirill Makarov 
Nebria (Epinebriola) christinae Huber & Schmidt, 2007 Paratype Male adult
Nepal, Rolwaling valley, Na to Yarlung Ri Base Camp, 4200–4900m, 23. 5. 2000, leg. J. Schmidt.
Copyright ©
Charles Huber 
Nebria (Epinebriola) molendai Huber & Schmidt, 2007 Paratype Male adult
Nepal, Rolwaling valley, Na to Omai Tsho lake, 4100–4500 m, 22. 5. 2000, leg. Schmidt
Copyright ©
Charles Huber 
Nebria (Nippononebria) kyushuensis Habu, 1958 Holotype Male adult
Mt. Raizan, FUKUOKA, 19. I. 1953 Coll. A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) kyushuensis Habu, 1958 (Head) Holotype Male adult
Mt. Raizan, FUKUOKA, 19. I. 1953 Coll. A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) kyushuensis Habu, 1958 (Pronotum) Holotype Male adult
Mt. Raizan, FUKUOKA, 19. I. 1953 Coll. A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) kyushuensis Habu, 1958 (Label) Holotype Male adult
Mt. Raizan, FUKUOKA, 19. I. 1953 Coll. A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) pusilla pusilla (as teres Habu, 1958) Holotype Male adult
Mt. Norikura, NAGANO, 04. VIII. 1952, N. Fukuhara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) pusilla teres Habu, 1958 (Lable) Holotype Male adult
Mt. Norikura, NAGANO, 04. VIII. 1952, N. Fukuhara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) pusilla pusilla (as teres Habu, 1958) Holotype Male adult
Mt. Norikura, NAGANO, 04. VIII. 1952, N. Fukuhara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nippononebria) pusilla pusilla (as teres Habu, 1958) Holotype Male adult
Mt. Norikura, NAGANO, 04. VIII. 1952, N. Fukuhara
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nebria) hikosana Habu, 1956 Holotype Male adult
Mt. Hiko, FUKUOKA, 23. V. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nebria) hikosana Habu, 1956e: 170 Holotype Male adult
Mt. Hiko, FUKUOKA, 23. V. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nebria) hikosana Habu, 1956e: 170 Holotype Male adult
Mt. Hiko, FUKUOKA, 23. V. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Nebria) hikosana Habu, 1956e: 170 Holotype Male adult
Mt. Hiko, FUKUOKA, 23. V. 1950, A. Habu
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Sadonebria) chinensis tsushimae Habu, 1981 Holotype Male adult
Nita, Tsushima Is., NAGASAKI, 1. V. 1975, K. Terada
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Sadonebria) chinensis tsushimae Habu, 1981 Holotype Male adult
Nita, Tsushima Is., NAGASAKI, 1. V. 1975, K. Terada
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Sadonebria) chinensis tsushimae Habu, 1981 Holotype Male adult
Nita, Tsushima Is., NAGASAKI, 1. V. 1975, K. Terada
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Sadonebria) chinensis tsushimae Habu, 1981 Holotype Male adult
Nita, Tsushima Is., NAGASAKI, 1. V. 1975, K. Terada
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Falcinebria) taketoi Habu, 1962a: 2 Holotype Male adult
Mikurigaike, Mt. Tateyama, TOYAMA, 5. VIII. 1961, A. Taketo<br />
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.<br />
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)<br />
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Falcinebria) taketoi Habu, 1962a: 2 Holotype Male adult
Mikurigaike, Mt. Tateyama, TOYAMA, 5. VIII. 1961, A. Taketo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Falcinebria) taketoi Habu, 1962a: 2 Holotype Male adult
Mikurigaike, Mt. Tateyama, TOYAMA, 5. VIII. 1961, A. Taketo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Falcinebria) taketoi Habu, 1962a: 2 Holotype Male adult
Mikurigaike, Mt. Tateyama, TOYAMA, 5. VIII. 1961, A. Taketo
Deposited in: National Institute for Agro-Environmental Sciences (NIAES), Tsukuba.
(http://www.niaes.affrc.go.jp/inventory/insect/dbcarabidae/carabidaetop.html)
Hiraku Yoshitake © Prohibited secondary use of all data
Copyright ©
Hiraku Yoshitake 
Nebria (Eunebria) limbigera limbigera Solsky, 1874: 12 Male adult
Kyrgyzstan, N Tien-Shan, Kyungey Ala-Too Mt. Rng, h =2500 m, 5.VII.2001, Dolin V.G. leg.
L=13.5 mm
Copyright ©
Ruslan Panin 
Leistus (Neoleistus) longipennis Casey, 1920
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Nakanebria) turmaduodecima Kavanaugh, 1981
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) piute piute s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) piute sevieri Kavanaugh, 1984
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) purpurata LeConte, 1878
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) schwarzi beverlianna Kavanaugh, 1979
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) schwarzi schwarzi s.str.
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) trifaria pasquineli Kavanaugh, 1984
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) trifaria trifaria s.str.
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) vandykei vandykei s.str.
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Catonebria) vandykei wyeast Kavanaugh, 1979
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Reductonebria) sahlbergi sahlbergi s.str.
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Reductonebria) sahlbergi triad Kavanaugh, 1979
Female adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Reductonebria) suturalis LeConte, 1850
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Reductonebria) zioni oasis Kavanaugh, 1979
Male adult
Mackenzie Flight, SI-EOL Intern; TL Erwin Lab, NMNH
Copyright © Mackenzie Flight
Nebria (Patrobonebria) himalayica Bates, 1889b: 212 Male adult
N INDIA, Uttarakhand, 13-15.04.2012, Uttarkashi distr.
N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi, 2300 m, Anichtchenko leg., det. & col.
Copyright ©
Alexander Anichtchenko 
Nebria (Sadonebria) sadona Bates, 1883: 217
Male adult
Collection ground Hatto, Tottori Pref. 35357<br />
(C) Working Group for Biological Indicator Ground Beetles Database, Japan<br />
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Nebria (Sadonebria) sadona Bates, 1883: 217
Male adult
Collection ground Hatto, Tottori Pref. 35357
(C) Working Group for Biological Indicator Ground Beetles Database, Japan
Prohibited secondary use of all data
Copyright © Katsuro Yahiro
Nebria (Epinebriola) oxyptera alzonai Deuve et Ledoux, 1989: 161 Holotype
Male adult
Copyright © Gianluigi Alzona
Nebria (Nippononebria) changbaiensis Kavanaugh & Liang, 2010 Male adult
China, Jilin Prov., Changbaishan Mountains, 1900-2000 m, 42,05° N, 128,06° E, 16.07.2012, R. Dudko leg.
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Roman Dudko 
Nebria (Catonebria) pektusanica Horvatovich, 1973: 19 Male adult
China, Jilin Prov., Changbaishan Mountains, 1900-2000 m, 42,05° N, 128,06° E, 16.07.2012, R. Dudko leg.
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Roman Dudko 
Nebria (Boreonebria) nivalis (Paykull, 1798) Male adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) hellwigii hellwigii Panzer, 1803 Male adult
NE Italy, Bolzano reg., Zirog, Mt. Schlüsseljoch, VII.1988, Manfed Egger leg.
L=13 mm
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Ruslan Panin 
Nebria (Alpaeonebria) germarii norica Schauberger, 1927e: 173 Male adult
W Austria, Tyrol, Mt. Kellerjoch h=2050m, VII.2004, Manfed Egger leg.
L=11 mm
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Ruslan Panin 
Nebria (Nebria) transsylvanica Germar, 1824 Male adult
W Ukraine, Transcarpathian Reg., Rakhov Distr., Chornogora Mt. Range, Mt. Gutyn-Tomnatyk h=2015m, alpin zone, 8.VI-27.VII.2011, Panin R. leg.
L=11 mm
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Ruslan Panin 
Nebria (Boreonebria) lacustris Casey, 1913 Lectotype
Female adult
Image from Kavanaugh et al, 2011
Copyright © ---
Nebria (Boreonebria) lacustris Casey, 1913 Lectotype
Female adult
Image from Kavanaugh et al, 2011
Copyright © ---
Nebria (Boreonebria) bellorum Kavanaugh, 1979
Male adult
Smokemont, Swain Co., North Carolina
Image from Kavanaugh et al, 2011
Copyright © ---
Nebria (Eunebria) jockischi jockischii Sturm, 1815 Male adult
SE France, Alpes-Maritimes, Mt. Col de Turini h=1500m, Graben SW, 18.VIII.2005, Manfred Egger leg.
L= 13 mm
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Ruslan Panin 
Nebria (Reductonebria) altaica Gebler, 1847 Male adult
Russia, Central Altay Mts., Kadrin riv., h = 845 m., 23.07.2000, А. Matalin leg.<br />
Фотография с сайта zin.ru\Animalia\Coleoptera. Автор - М.Э. Смирнов
Copyright ©
Maksim Smirnov 
Nebria (Tyrrhenia) holtzi K. Daniel, 1903a: 162 Female adult
Greece, N-Peloponnese, Kalentzi vill., Mt. Enimanthos, 11.VI.2006, Manfred Egger leg.
L=13 mm
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Ruslan Panin 
Leistus (Leistus) piceus piceus Frolich, 1799 Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 9mm
Copyright ©
Aleš Sedláček 
Leistus (Leistus) nitidus (Duftschmid, 1812) Male adult
SW Romania, Munţii Retezat; 8mm
Copyright ©
Aleš Sedláček 
Leistus (Pogonophorus) montanus kultianus Farkac & Fassati, 1999 Female adult
CZ, N Bohemia, České středohoří; 8mm
Copyright ©
Aleš Sedláček 
Leistus (Pogonophorus) montanus pawlowskii Farkac & Fassati, 1999 Male adult
S Poland, Babia Góra Mt.; 7mm
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Aleš Sedláček 
Leistus (Pogonophorus) montanus corconticus Farkac & Fassati, 1999 Male adult
N Bohemia, Krkonoše Mts.; 7mm
Copyright ©
Aleš Sedláček 
Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843 Male adult
SW Bosnia, Bijakoviči - Križevac Mt.; 8mm
Copyright ©
Aleš Sedláček 
Nebria (Nebria) kratteri pindica Jeanne, 1974 Male adult
C Greece, Thessalia, Kastraki; 10mm
Copyright ©
Aleš Sedláček 
Nebria (Alpaeonebria) fasciatopunctata L. Miller, 1850: 12 Female adult
NE Slovenia, Pohorje Mts., Ruše; 10mm
Copyright ©
Aleš Sedláček 
Nebria (Nebria) tatrica tatrica L. Miller, 1859: 304 Female adult
SK, N Slovakia, Vysoké Tatry Mts.; 10mm
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Aleš Sedláček 
Nebria (Eunebria) picicornis picicornis Fabricius, 1801 Male adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 13mm
Copyright ©
Aleš Sedláček 
Nebria (Eunebria) jockischii hoepfneri Dejean, 1826 Female adult
CZ, NE Moravia, Moravskoslezské Beskydy Mts.; 12mm
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Aleš Sedláček 
Nebria (Nebria) brevicollis Fabricius, 1792 Male adult
SK, NW Slovakia, Malá Fatra Mts.; 12mm
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Aleš Sedláček 
Nebria (Nebria) dahlii dahilii Duftschmid, 1812: 50 Male adult
NW Slovenia, Trnovski Gozd Mts.; 8mm
Copyright ©
Aleš Sedláček 
Nebria (Alpaeonebria) fasciatopunctata L. Miller, 1850: 12 Male adult
NE Slovenia, Pohorje Mts., Ruše; 9mm
Copyright ©
Aleš Sedláček 
Nebria (Alpaeonebria) fuscipes Fuss, 1850 Female adult
SK, NE Slovakia, Vihorlat; 13mm
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Aleš Sedláček 
Nebria (Paranebria) livida livida Linne, 1758 Female adult
CZ, N Moravia, Choryně; 14mm
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Aleš Sedláček 
Nebria (Boreonebria) gyllenhali gyllenhali Schonherr, 1806 Female adult
CZ, NE Moravia, Staré Hamry; 9mm
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Aleš Sedláček 
Nebria (Nebria) salina Fairmaire et Laboulbene, 1854: 14 Male adult
CE Germany, Dresden; 11mm
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Aleš Sedláček 
Nebria (Nebria) tatrica dumbirensis Pulpan, 1957: 121 Female adult
SK, C Slovakia, Nízké Tatry Mts.; 9mm
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Aleš Sedláček 
Nebria (Nebria) tatrica fatrensis Hurka et Pulpan, 1992 Female adult
SK, NW Slovakia, Malá Fatra Mts.; 12mm
Copyright ©
Aleš Sedláček 
Nebria (Nebria) transsylvanica Germar, 1824 Male adult
C Romania, E Carpathiansю, Baiu Mts., Traisteni; 11mm
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Aleš Sedláček 
Nebria (Oreonebria) diaphana bohiniensis J. Muller, 1926 Female adult
NW Slovenia, Trnovski Gozd Mts.; 6,5mm
Copyright ©
Aleš Sedláček 
Nebria (Asionebria) mentoincisa Huber & Schmidt, 2013 Paratype Male adult
Tibet (South Central) 29.6.2007 / Dulong, Kurum vall., NW Lhasa / 4900–5200 m, leg. Schmidt / ca. 29°42’18 N, 90°35’16 E // south ascent of / Tsubu side valley / above Tsurphu Monastery
image from original describtion
Copyright ©
Charles Huber 
Leistus (Pogonophorus) lopatini Kabak, 2013 Paratype Male adult
Musart, China’s possession, 7000’ VI.915, Rückbeil
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Ilya Kabak 
Nebria (Nebriola) gosteliae Huber, 2010 Paratype Male adult
Copyright ©
Gianni Allegro 
Nebria (Nebria) asturiensis Bruneau de Mire, 1964: 31 Male adult
NW-Spain, Province Palencia, Cantabrian Mts, P.N. de Fuentes Carrionas, Puerto de Piedrasluengas pass, h=1329m, 14.V.2003, T.Gazurek leg.
L=11.5 mm
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Ruslan Panin 
Nebria (Nakanebria) shibanai shiretokoana Nakane, 1960: 64 Male adult
Copyright ©
Kirill Makarov 
Nebria (Nakanebria) shibanai shiretokoana Nakane, 1960: 64 Female adult
Kunashir Island, right bord of valley of Severjanka River 44°20’09”N 146°01’16”E - 44°19’51”N 146°01’43”E, h~115 29.VI.2008 leg. K.Makarov
Copyright ©
Kirill Makarov 
Leistus (Leistus) frater Reitter, 1897c: 121 Male adult
det. K. Makarov
Copyright ©
Kirill Makarov 
Nebria (Catonebria) praedicta Kavanaugh & Schoville, 2009 Male adult
USA, CA: Trinity Co, Thompson Peak, 2150-2500 m, 41°01' N, 123°03' W, 16.05.2013, R. Dudko, S. Schoville leg.
Copyright ©
Roman Dudko 
Leistus (Leistus) semenovi Perrault, 1986 Female adult
Copyright ©
Kirill Makarov 
Nebria (Patrobonebria) capillosa Ledoux et Roux, 1992d: 142 Male adult
N INDIA, Uttarakhand, 13-15.04.2012, Uttarkashi distr.
N30°57'41.57", E78°41'54.75" left tributary of Bhagirathi, 2300 m, Anichtchenko leg., det. & col.
Copyright ©
Alexander Anichtchenko 
Nebria (Eonebria) kurentzovi Lafer, 1989 Female adult
Maritime prov., Shkotovo Distr., Livadijsky Mt.R., Litovka (Falaza) Mt.,h~850m 30.VII.2014 leg. S.Ivanov
L=13.3 mm
Copyright ©
Kirill Makarov 
Nebria (Catonebria) sajana sarlyk Dudko et Shilenkov, 2001 Male adult
Altai, Seminskiy Mt.R., Mt. Sarlyk, h~1950-2250 13.VI.2013 leg. R. et E. Dudko
Copyright ©
Kirill Makarov 
Nebria (Catonebria) sajana sarlyk Dudko et Shilenkov, 2001 Female adult
Altai, Seminskiy Mt.R., Mt. Sarlyk, h~1950-2250 13.VI.2013 leg. R. et E. Dudko
Copyright ©
Kirill Makarov 
Leistus (Leistus) piceus piceus Frolich, 1799 Female adult
Copyright ©
Kirill Makarov 
Nebria (Boreonebria) subdilatata Motschulsky, 1844 Male adult
det. K. Makarov
Copyright ©
Kirill Makarov 
Leistus (Pogonophorus) spinangulus Reiter, 1913 Male adult
China, Xinjiang, Narat Mt. R., Dzhergalan Riv., forest, 42°56′31″N / 82°34′09″E, 2645 m, 27.07.2013, I.I. Kabak leg.
Copyright ©
Ilya Kabak 
Leistus (Pogonophorus) spinangulus Reiter, 1913 Female adult
China, Xinjiang, Narat Mt. R., Dzhergalan Riv., forest, 42°56′31″N / 82°34′09″E, 2645 m, 27.07.2013, I.I. Kabak leg.
Copyright ©
Ilya Kabak 
Leistus (Pogonophorus) spinangulus Reiter, 1913 Lectotype Female adult
(HNHM), «Kuldscha», «Monotypus 1913 Leistus (Euleistulus) spinangulus Reitter», «L. spinangulus mi 1913 Type», «Lectotypus Leistus spinangulus Reitt. des. Shilenkov, 1986», «coll. Reitter».
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Ilya Kabak 
Leistus (Pogonophorus) kazenasi Kabak, 2015 Paratype Male adult
China, Xinjiang Prov., Narat Mt. R., SSW of Tekes Vill., Kokterek bass., Buryl Riv., 3145 m, 42°48′06″N / 81°41′20″E, 28.07.2009, I.I. Kabak leg.
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Ilya Kabak 
Leistus (Pogonophorus) kazenasi Kabak, 2015 Paratype Female adult
China, Xinjiang, Koeksu Vall., Dunsala Riv., S of Tekes, 42°42′42″N / 81°51′12″E, h = 2655 m, 7.07.2012, I.I. Kabak leg.
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Ilya Kabak 
Nebria (Eunebria) psammodes (P. Rossi, 1792) Male adult
C-Italy, Toscana, Pistoia (PT), Pracchia env.,16.IX.1971, Campadelli leg.
L= 13 mm
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Ruslan Panin 
Nebria (Nebria) tibialis Bonelli, 1810: 54 Female adult
Italy, near Florence, Campigna National Park h~1200m, 17.VII.1969, P.Garagnani leg.
L=13 mm
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Ruslan Panin 
Nebria (Nebriola) cordicollis escheri Heer, 1837: 17 Male adult
N Italy, Lombardia,(SO), Valmalenco , Glacier del Ventina, 23.VII.1997, L. Diotti leg.
L=10 mm
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Ruslan Panin 
Nebria (Oreonebria) castanea castanea Bonelli, 1810 Male adult
NW Italy, Piemonte, Cuneo prov., Crissolo env., 17.VI.1989, L. Diotti legю
L=10 mm
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Ruslan Panin 
Leistus (Leistus) fulvus Chaudoir, 1846a: 105 Male adult
Copyright ©
Kirill Makarov 
Leistus (Pogonophorus) rufomarginatus (Duftschmid, 1812) Male adult
Copyright ©
Kirill Makarov 
Leistus (Pogonophorus) alaiensis Kabak, 1995: 9 Female adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) cf. retrospinosa Heyden, 1885 Female adult
det. Anichtchenko A. 2018
Copyright ©
Kirill Makarov 
Nebria (Alpaeonebria) crenatostriata Bassi, 1834 Female adult
NW Italy, Vercelli prov., Alagna Valsesia, A.Zube h~2400 m, 14.VIII.1979, R. Monguzzi leg.
L=14 mm
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Ruslan Panin 
Nebria (Nakanebria) shibanaii shibanaii Ueno, 1955d: 45 Male adult
Japan,Hokkaido, Mt. Asahi dake 26.VII.1975, S.Morita leg.
Copyright ©
Kirill Makarov 
Nebria (Nakanebria) shibanaii shibanaii Ueno, 1955d: 45 Female adult
Japan, Hokkaido, Mt. Daisetsuzan, 3.VII.1955, Kawamura leg.
Copyright ©
Kirill Makarov 
Leistus (Pogonophorus) depressus depressus Male adult
Kyrghyzstan, E part of Kirghizskyi Mt.R., Keghety River
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- - - 
Leistus (Pogonophorus) rousi Pulpan et Reska, 1977 Holotype Male adult
Image from types-nmp.cz
Copyright ©
- - - 
Nebria (Eunebria) kazemii Wrase & Schnitter, 2017 Paratype Female adult
IRAN (Kerman Prov.) / waterfall at Darb-e Āsīāb vill. / nr. Kuhpayeh vill. N Kerman / 2600 m / N 30°31'18.4"/E 57°09'44.6" (canyon, in gravel/under stones) / 7.VI.2014 D.W. Wrase & B. Laser. Image from Wrase & Schnitter, 2017
Copyright ©
- - - 
Nebria (Eunebria) boiteli Alluaud, 1932 Male adult
Moyen Atlas, Lac Sidi Ali. Image from Wrase & Schnitter, 2017
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- - - 
Archileistobrius hwangtienyuni Shilenkov et Kryzhanovskij, 1983 Female adult
China, Sichuan, Emeishan, 160km SSW Chengdu, 1700m, 22.6.1994, leg. Schillhammer; 10mm (Collection R. Sciaky)
Copyright ©
Alexander Anichtchenko 
Archastes cf. solitarius Ledoux et Roux, 1999 Male adult
China, Shanxi prov., Zhouzhi Co., ouzhenzi env., 3200m, 17-23.7.1998
Copyright ©
Alexander Anichtchenko 
Leistus (Pogonophorus) ovipennis Chaudoir, 1867: 262 Male adult
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Vittorino Monzini 
Nebria (Nebria) kratteri kratteri Dejean, 1831 Female adult
S Italy, Calabria, Aspromonte Massif, VII.1984, E.Monzini leg., L=15 mm
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Ruslan Panin 
Nebria (Nebria) azarbayanei Muilwijk, 2015 Holotype Male adult
Iran, Azarbayjan-e Gharbi/ Khoy – Siyah Chesmeh road/17 km W Zar Abad 2640m/N 38’45’39” E 044’28’35”/30.08.2008, lg. Frisch & Serri
Copyright ©
Jan Muilwijk 
Nebria (Eunebria) ferganensis urumbashi Kabak et Putchkov, 1996: 115 Paratype Male adult
W Kyrgyzstan, In.Tien-Shan, Fergana Mt Range, Urumbash riv., 40 km W Kazarman, h~2500 m, A.Puchkov leg.
L=12 mm
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Ruslan Panin 
Nebria (Orientonebria) coreica Solsky, 1875 Male adult
Copyright ©
Kirill Makarov 
Leistus (Leistus) megrelicus Shilenkov, 1999 Paratype Male adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) fischeri Faldermann, 1836a: 31 Female adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) verticalis Fischer von Waldheim, 1828: 249 Female adult
Copyright ©
Kirill Makarov 
Leistus (Leistus) denticollis Reitter, 1887 Female adult
Russia, Krasnodar reg., Achishkha mt., 1500 m, 4.8.1993
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Alexander Anichtchenko 
Leistus (Leistus) elegans Rost, 1891 Female adult
C Caucasus, N Ossetia, Urukh riv., 2300 m, 23.7.1997, A. Koval leg.
Copyright ©
Alexander Anichtchenko 
Leistus (Pogonophorus) spinibarbis rufipes Chaudoir, 1843 Female adult
Cyprus, Larnaca reg., Valva
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Alexander Anichtchenko 
Leistus (Pogonophorus) expansus Putzeys, 1874 Male adult
Spain, Sierra de la Solana de Alcudia, pto Pulido, 850 m (Ciudad Real) 17.11.2005, A. Anichtchenko leg. & det.
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Alexander Anichtchenko 
Leistus (Sardoleistus) sardous Baudi di Selve, 1883 Male adult
Italy, Toscana, Migliarino, (PI), 28.4.1972
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Alexander Anichtchenko 
Leistus (Leistus) caucasicus Chaudoir, 1867 Male adult
Libano, Zghorta prov, Horsh Ehden, 13.5.2000
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Alexander Anichtchenko 
Leistus (Leistus) semenovi Perrault, 1986 Male adult
SE Kazakhstan, Zailijsky mt rng., Bol Almaatinka riv., 3000 m 17.6.2000, A. Klimenko leg.
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Alexander Anichtchenko 
Leistus (Pogonophorus) tschitscherini Semenov, 1906a: 3 Male adult
SE Kazakhstan, C Tian Shan, Bajankol riv, Bajankol vill. 3000m 1.7.2000, A. Klimenko leg.
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Alexander Anichtchenko 
Leistus (Leistus) fulvibarbis Dejean, 1826 Female adult
Spain, Sierra Nevada, rio Lanjaron, Cortijo de Ballesteros, 1800m, 29-31.3.2010, Anichtchenko A. leg. & det.
Copyright ©
Alexander Anichtchenko 
Leistus (Leistus) fulvibarbis Dejean, 1826 Male adult
Greece, Ahaia, M. Erimantos, 1100m Kalentzi, 3.5.2004, F. Angelini leg. det. Anichtchenko A.
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Alexander Anichtchenko 
Leistus (Leistus) angusticollis Dejean, 1826 Male adult
Spain, Avila, El Tiemblo, 1600 m , 18.11.2007, Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Leistus (Leistus) oopterus Chaudoir, 1861 Male adult
Spain, Sierra de la Cabrera Baja, 8 km SW Truchillas, 9.7.2008, Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Leistus (Leistus) valcarceli Wrase, Riuz-Tapiador & Zaballos, 1998 Male adult
Spain, Sierra de la Cabrera Baja, 1 km W Pena Negra Mt., 2000m 9.7.2008, Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Leistus (Leistus) constrictus Schaufuss, 1862 Male adult
Spain, Sierra Guadarrama,, Montes Carpetinos, Pto. De Somosierra, 1600-1700m, 3.6.2006 Anichtchenko A.
Copyright ©
Alexander Anichtchenko 
Nebria (Paranebria) livida livida Linne, 1758 Female adult
Copyright ©
Kirill Makarov 
Nebria (Paranebria) livida livida Linne, 1758 Female adult
Copyright ©
Kirill Makarov 
Nebria (Pseudonebriola) dabanensis Shilenkov 1982 (= sajanica) Female adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) tristicula Reitter, 1888 Male adult
? tenella
Copyright ©
Kirill Makarov 
Nebria (Pseudonebriola) kaszabi Shilenkov, 1982 Male adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) elbursiaca B. Bodemeyer, 1927 Male adult
Iran, 40 km S Ardabil, Goli, 20.5.2005 Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Nebria (Reductonebria) angustula Motschulsky, 1866 Female adult
Russia, Kamchatka, Vilyuchinskii volcan vic 900 m, alpic, 29.7.1990, S. Saluk leg. 13 mm
Copyright ©
Alexander Anichtchenko 
Leistus (Neoleistus) ferruginosus Mannerheim, 1843 Male adult
USA, Wa, Colwitz Co., Woodland 10 mi E, 21.7.2012
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Alexander Anichtchenko 
Nebria (Boreonebria) baicalica Motschulsky, 1844 Male adult
Russia, E Siberia, Listvianka vill., lake Baikal shore, 25.8.1992, Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Nebria (Pseudonebriola) saurica Shilenlkov, 1976 Male adult
NE Kazakhstan, Saur mt. rng., Sargar Mt., 12-15.6.2000, S.Nekrashevich leg.
Copyright ©
Alexander Anichtchenko 
Nebria (Boreonebria) suvorovi Shilenkov, 1976 Female adult
Kirgizstan, Kungei Ala Too, Kichi-Urukty v., 2800 m, 1.7.1999
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Alexander Anichtchenko 
Nebria (Nebria) atlantica Oberthur, 1883 Male adult
Morocco, Grand Atlas Mts., 2200-2300 m, Oukaimeden env., 16-17.6.2010, Anichtchenko A. leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Nebria) andalusia Rambur, 1837 Male adult
Spain, Sevilla, Cerro de Hierro, 18.1.2003, Anichtchenko A. leg. & det.
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Alexander Anichtchenko 
Nebria (Sadonebria) chinensis Bates, 1872 Male adult
China, Liaoning prov., Kuandian co., Banlaing, Ba-che-chuan 6.2006 13 mm
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Alexander Anichtchenko 
Leistus (Evanoleistus) pseudocrenifer Sciaky, 1995 Male adult
China, Sichuan, Min Shan, Baima pass, 3500 m, 6.7.2004. det. R. Sciaky, 2018
Copyright ©
Alexander Anichtchenko 
Nebria (Boreonebria) biseriata Lutshnik, 1915 Male adult
Russia, Khabarovsk reg., Bereinskii Nat. res.
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Alexander Anichtchenko 
Nebria (Nebria) hemprichi Klug, 1832 Male adult
Iran, Lorestan, Khorram Abad, S of Mahmudvand, 30.4.2007 Anichtchenko A. leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Eonebria) komarovi Semenov & Znojko, 1928 Male adult
Russia, Primorie reg., Chernye Gory Mts. det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Reductonebria) ochotica R.F. Sahlberg, 1844 Female adult
Russia, Khabarovsk reg., Verhnebureinskii diistr., Bureya riv vall., Ussomah riv. 22.7.2006
Copyright ©
Alexander Anichtchenko 
Nebria (Nebria) asturiensis Bruneau de Mire, 1964 Male adult
Spain, Peña Ubiña, 2000 m, 28.6.2009, Anichtchenko A. leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Boreonebria) pazi Seidlitz, 1867 Male adult
Spain, Gredos Mts., 2 km E from La Mira Mt., 4-7.7.07, 2000 m, A. Anichtchenko leg. & det.
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Alexander Anichtchenko 
Nebria (Nebria) sobrina Schaufuss, 1862 Female adult
Spain, Mts. Cantabricos, Soto de Sajambre, Canto Cabronero mt., 1850 m, 12.8.2008, A. Anichtchenko leg. & det.
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Alexander Anichtchenko 
Nebria (Catonebria) sajana sarlyk Dudko et Shilenkov, 2001 Male adult
Russia, C Altay Mts., Seminsky mt. rng., Sarlyk mt., 1900m, 19.6.2002 Menshikov A leg. ; det. Anichtchenko A.
Copyright ©
Alexander Anichtchenko 
Nebria (Boreonebria) bargusinica Shilenkov, 1999 Male adult
N Baikal lake, Baikalskii mt. rng., Izumrudnoe lake, 6.7.2013. det. Anichtchenko A. 2018
Copyright ©
Alexander Anichtchenko 
Nebria (Eunebria) jockischii bolivari Jeanne, 1966 Female adult
Spain, Sierra Nevada, Pradollano, 2.6.2005, rio Monachil, 2000 m., A.Anichtchenko leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Nebria) lafresnayei lafresnayei Audinet-Serville, 1821 Male adult
Spain, Pirineos, Canfranc, Cabana de la vuelta de Iserias, 1800 m, 16.6.2005, A. Anichtchenko leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Nebria) lafresnayei lafresnayei Audinet-Serville, 1821 Male adult
Spain, Pirineos, Canfranc, Cabana de la vuelta de Iserias, 1800 m, 16.6.2005, A. Anichtchenko leg. & det.
Copyright ©
Alexander Anichtchenko 
Nebria (Alpaeonebria) rhilensis J. Frivaldszky, 1879 Male adult
Bulgaria Rodopi, Cernatica-Goljam, Persenk 1400-1900m.
Copyright ©
Alexander Anichtchenko 
Nebria (Nebria) bonellii M.F. Adams, 1817 Female adult
Copyright ©
Kirill Makarov 
Nebria (Catonebria) mellyi teletskiana Dudko et Shilenkov, 2001 Female adult
Copyright ©
Kirill Makarov 
Nebria (Eunebria) talassica Shilenkov, 1982 Male adult
Copyright ©
Kirill Makarov 
Nebria (Eunebria) tschatkalica Kabak & Shilenkov, 2001 Male adult
Copyright ©
Kirill Makarov 
Nebria (Nebria) fischeri Faldermann, 1836 Female adult
N Turkey, Trabzon env. 26.7.1996 det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Nebria) patruelis Chaudoir, 1846 Male adult
Georgia, Tehuri riv., 2000 m, 9.8.1990; det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Nebria) posthuma K. Daniel et J. Daniel, 1891 Male adult
Italy, Abruzzo Gran Sasso, Campo Imperat., rifugio, 2100m 5,7.2000, F. Angelini leg.
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Alexander Anichtchenko 
Nebria (Nebria) retrospinosa Heyden, 1885 Female adult
Karachaevo-Cherkessia, Beskes riv., 2300 m, 12.8.97 Solodovnikov I. leg.; det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Nebria) tenella Motschulsky, 1850 Male adult
Karachaevo-Cherkessia, Pjiya vill vic., Abishira-Akhuba Mt.R., Bolshaya Laba riv. 16.7.1989; det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Nebria) tristicula Reitter, 1888 Female adult
Caucasus, Adygea, Lago Naki plato, 3.8.2004; det. Anichtchenko A. 2018
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Alexander Anichtchenko 
Nebria (Nebria) bonellii M.F. Adams, 1817 Male adult
NW Caucas, Adygea reg., Lago Naki plato, Murzikal mt., 2100 m, 28.6.07
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Alexander Anichtchenko 
Nebria (Nebria) heinziana Shilenkov, 1983 Male adult
N Turkey, Trabzon env., 26.7.1996
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Alexander Anichtchenko 
Leistus (Evanoleistus) championi Andrewes, 1920 Male adult
India, Uttarakhand, 14 km E Uttarkashi, leg. Anichtchenko A.; det. J. Schmidt, 2018
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Alexander Anichtchenko 
Nebria (Tyrrhenia) vanvolxemi Putzeys, 1874 Female adult
Portugal, Baragem dos pisoes sierra de Monchique, Algarve, 20-09-1983
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Pascal Deschamps 
Nebria (Tyrrhenia) testacea Olivier, 1811 Male adult
Grèce, Crète, Heraklion, Lentas, 06-12-2007
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Pascal Deschamps 
Nebria (Barbonebriola) rubostipes Huber & Schmidt, 2017 Holotype Male adult
Nepal, Prov. Karnali, Distr. Humla, 16 km W Simikot, 2 km NW Sankha La, 4250–4950 m, 29°56'39" N, 81°39'02" E, 29.VI.2001, leg. E. Grill
(NME). Scale bar 5mm
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Joachim Schmidt 
Nebria (Barbonebriola) tenuisulcata Huber & Schmidt, 2017 Holotype Female adult
Nepal, Mahakali/Darchula, 12 km N Ghusa, high valley SSW Api, 4100 m, 29°56'28" N, 80°54'24" E, stone debris, 8.VI.2005, leg. A. Weigel (NME). Scale bar 5mm
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Joachim Schmidt 
Nebria (Barbonebriola) kagmara Huber & Schmidt, 2017 Paratype Male adult
Nepal, Prov. Karnali, Distr. Dolpo, Kagmara La, westside, High Camp,
4300 m, 29. 6. 2009, leg. Santos Tamang (NME).
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Joachim Schmidt 
Nebria (Epinebriola) triseriata Huber & Schmidt, 2017 Paratype Male adult
Nepal, Dhaulagiri Zone, Myagdi Distr., Annapurna South Himal, W-Slope, 28° 28,30' N, 83° 44,55' E, 4500 m, 22. 5. 2001, leg. G. Hirthe & J. Schmidt (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) rupina Huber & Schmidt, 2017 Paratype Male adult
Nepal, Manaslu Himal, Rumche Tal to Rupina La pass, 4200 mNN, 24. 6. 2001, leg. S. Tamang (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) montisanimae Huber & Schmidt, 2017 Paratype Male adult
Nepal, Manaslu Himal, Narte Pokhari, S of Rupina La pass, 4400 m, 26. 6. 2001, leg. S. Tamang (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) impunctata Huber & Schmidt, 2017 Paratype Male adult
Nepal, Ganesh Himal, Jaisuli Kund env., 4300–4500 m, 13.–16.6. 2000, leg. Iman Ghalé, Santos Tamang, Ram, Santa & Santé Gurung (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) tuberculata Huber & Schmidt, 2017 Holotype Female adult
Nepal, Helambu, S-slope Ganja La, 4400–4800 m, 5. 8. 1998, leg. C. Berndt (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) numburica Huber & Schmidt, 2017 Holotype Male adult
Nepal, Solu Khumbu, S Dudh Kund, 4400–4600 m, 27°42' N, 86°35' E, 25. V. 2013, leg. J. Schmidt (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) retingensis Huber & Schmidt, 2017 Holotype Male adult
Tibet, side valley S Reting, 30°15'57" N, 91°32'04" E, 4500–4800 m, 1.8.2015, leg. J. Schmidt (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) delicata Huber & Schmidt, 2017 Holotype Male adult
S Tibet, Mt. Mendju Zari, SE Lhasa, 29°34' N, 91°13' E, 5200– 5420 m, 4.6.2011, leg. J. Schmidt (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) pseudorestias Huber & Schmidt, 2017 Holotype Male adult
Nepal, Jaljale Himal, Thangla Bhanjyang, 4630 m, 29.VI.2011, leg. S. Tamang, 27°41'41" N, 87°35'32" E (cSCHM).
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Joachim Schmidt 
Nebria (Epinebriola) incarinata Huber & Schmidt, 2017 Holotype Male adult
India, Arunachal Pradesh, Sela Pass, 27°30'12" N, 92°06'16" E, 4200–4500 m, 24./25. 6. 2008, leg. Reuter (cSCHM).
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Joachim Schmidt 
Leistus (Evanoleistus) colpodes Deuve, 2018 Holotype Female adult
Chine, Sichuan, Siguniang Shan, Rilong, 3800- 4300 mètres, 31°02’58’’N-102°16’85’’E, in coll. M. Janata, à Prague (République Tchèque)
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Thierry Deuve 
Nebria (Catonebria) splendida Fischer von Waldheim, 1844 Male adult
Kazakhstan, Taldi Kurgan, 30-06-1990
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Pascal Deschamps 
Nebria (Nebriola) laticollis Dejean, 1826: 244 Female adult
France 05350 Chateau Ville Vieille Bois Noir 18-07-2014
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Pascal Deschamps 
Nebria (Oreonebria) macrodera K. Daniel, 1903 Female adult
France, 06430 Tende col de Tende, 19-06-2016
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Pascal Deschamps 
Leistus (Leistus) barnevillei Chaudoir, 1867 Female adult
SPAIN, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg.; det. Anichtchenko A., 2019
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Alexander Anichtchenko 
Leistus (Leistus) acutangulus Perrault, 1979 Male adult
Serra de Estella, Manteigas, Guarda, Portugal, 20.7.1986 M. Toribio leg.
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Alexander Anichtchenko 
Leistus (Leistus) angulatus Piochard de la Brulerie, 1872 Male adult
Spain, Villoslada de Cameros, Logrono, 3.6.1987, M. Toribio leg.
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Alexander Anichtchenko 
Leistus (Leistus) lenkoranus Reitter, 1885d: 217 Male adult
S Azerbayddzan, Lenkoranskyi distr., Vazha riv., 2 km NW Dashdatuk vill., 17.4.2004, K. Makarov leg.
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Alexander Anichtchenko 
Leistus (Pogonophorus) terskeiensis Belousov et Kabak, 1992 Female adult
Terskey Alatoo, Barskaun 15.8.84 Ovchinnikov leg. det. K. Makarov, 2019
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Alexander Anichtchenko 
Nebria (Nebria) andarensis Bolivar y Pieltain, 1923 Male adult
Spain, Mt. Cantabricos, Picos de Europa, 2200 m, 2 km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg.
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Alexander Anichtchenko 
Nebria (Nebria) andarensis Bolivar y Pieltain, 1923 Female adult
Spain, Mt. Cantabricos, Picos de Europa, 2200 m, 2 km NW mirador Fuente De, 22.07.2019, A. Anichtchenko & R. Sciaky leg.
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Alexander Anichtchenko 
Nebria (Eunebria) jockischii orensis Breit, 1914 Male adult
Spain, Cantabria, 2 km E Puerto de San Glorio, near Vejo riv., 43.072369, -4.733061, 1300 m., 23.07.2019, A. Anichtchenko & R. Sciaky leg. det. Anichtchenko A.
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Alexander Anichtchenko 
Nebria (Falcinebria) niohozana Bates, 1883 Lectotype Male adult
Image from Sasakawa, 2020
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Nebria (Falcinebria) reflexa Bates, 1883 Lectotype Male adult
Image from Sasakawa, 2020
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Nebria (Catonebria) baumanni Kavanaugh, 2015 Holotype Male adult
Copyright ©
David Kavanaugh 
Leistus (Nebrileistus) kratkyi Machado, 2020 Holotype Male adult
Spain, Canary Islands, La Gomera, El Cedro, 28°07’48” N, 17°13’06” W, 31 Oct. 2018, V. Dušánek leg., genitalia embedded in alcohol-polyvinyl on plastic card
(TFMC//CO-16044). Image from Machado, 2020
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Leistus (Nebrileistus) nubivagus Wollaston, 1864 Male adult
Image from Machado, 2020
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Leistus (Leistus) niger Gebler, 1847 Male adult
Far East, Lazovskij Nat. Res., Proselochnaja Riv. near nouth 43°00'34"N 134°07'43"E 10.VII.2005 K.Makarov leg.
L=11.0 mm
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Kirill Makarov 
Cicindis horni Bruch, 1908
Archaeocindis johnbeckeri (Banninger, 1927) Holotype
Archaeocindis hormozensis Azadbakhsh, 2020 Paratype
Notiokasis chaudoiri Kavanaugh and Negre, 1982
Notiophilus aeneus (Herbst, 1806)
Notiophilus anichtchenkoi Barsevskis, 2009 Holotype
Notiophilus aquaticus Linne, 1758
Notiophilus biguttatus Fabricius, 1779
Notiophilus biguttatus Fabricius, 1779
Notiophilus borealis Harris, 1869
Notiophilus breviusculus Solsky, 1873
Notiophilus intermedius Lindroth, 1955
Notiophilus nemoralis Fall, 1906
Notiophilus novemstriatus LeConte, 1849
Notiophilus palustris Duftschmid, 1812
Notiophilus reitteri Spaeth, 1900
Notiophilus semistriatus LeConte, 1857
Notiophilus palustris (Duftschmid, 1812)
Notiophilus biguttatus Fabricius, 1779: 222 (Elaphrus)
Notiophilus dostali Barsevskis, 2011 Holotype
Notiophilus nuristanensis Barsevskis, 2011 Holotype
Notiophilus persicus Breit, 1914b: 162
Notiophilus ghilarovi Kryzhanovskij, 1995: 267
Notiophilus chihuahuae Casey, 1913
Notiophilus hiemalis Semenov et Arnoldi, 1937: 222 Holotype
Notiophilus sibiricus Motschulsky, 1844: 85 Holotype
Notiophilus rufipes Curtis, 1829: pl. 254
Notiophilus altaicus Kabak, 2014 Paratype
Opisthius richardsony Kirby, 1837
Paropisthius davidis Fairmaire, 1887
Paropisthius indicus indicus Chaudoir, 1863
Paropisthius masuzoi Kasahara, 1989
Paropisthius indicus chinensis Bousquet et Smetana, 1996: 225 Paratype
Pelophila borealis Paykull, 1790
Pelophila borealis Paykull, 1790
Pelophila borealis Paykull, 1790
Pelophila borealis Paykull, 1790
Pelophila rudis LeConte, 1863
