Gigadema obscurum Sloane, 1914
Male adult
Australia, Queensland, Brisbane. Image from: Mal, N., Drumont, A., Kerkhof, S. & P. Grootaert, 2012. Pictorial beetle collection of the Royal Belgian Institute of Natural Sciences. det. Anichtchenko A. 2020
Copyright ©

Helluodema brunneum Sloane, 1917 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema minutum Laporte, 1867 Holotype
Male adult

Copyright © Museums Victoria

Helluapterus niger Sloane, 1914 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema biordinatum Sloane, 1914 Syntypes
Male adult

Copyright © Museums Victoria

Gigadema mandibulare Blackburn, 1892 Syntypes
Male adult

Copyright © Museums Victoria

Helluo insignis Sloane, 1890 Paratype
Male adult

Copyright © - - -

Neohelluo angulicollis Sloane, 1914
Male adult
Australia. 24 mm
Copyright © Mark Golding

Dicranoglossus resplendens (Castelnau, 1867)
Male adult
Australia, Quennsl. Cloncurry, III.1993; 11mm (Collection R. Sciaky)
Copyright © Alexander Anichtchenko

Aenigma iris Newman, 1836
Female adult
Australia, Queensland, M. Garnet, 26.II.1992.
Copyright © Riccardo Sciaky

Helluosoma cf. bouchardi Baehr, 2005
Male adult
Australia (coll. R. Sciaky)
Copyright © Riccardo Sciaky

Helluonidius cyanipennis (Hope, 1842) Holotype
Male adult

Copyright © - - -

Ametroglossus ater (W.J.Macleay, 1887)
Male adult

Copyright © Creative Commons

Helluosoma atrum Castelnau, 1867
Male adult

Copyright © Creative Commons

Helluodema unicolor (Hope, 1842)
Male adult
Australia
Copyright © Creative Commons

Helluonidius aterrimus (W.J.Macleay, 1873)
Male adult

Copyright © Creative Commons

Epimicodema mastersii W.J.Macleay, 1871
Male adult

Copyright © - - -

Helluosoma bisetosum Baehr, 2017 Holotype
Female adult
“Ourimbah N. S. Wales J. Armstrong” (ANIC) 10.9 mm
Copyright © Martin Baehr

Helluosoma kalumburu Baehr, 2017 Holotype
Female adult
“14.17S 126.39E Kalumburu, W.A. 17 June 1985, L.A.Crawen” (ANIC).
Copyright © Martin Baehr

Helluosoma crenulicolle Baehr, 2017 Holotype
Male adult
“12.17S 133.20E Cooper Creek, N.T. 11km S.by W. of Nimbuwah Rock, 3.vi.73, Feehan & Upton / Helluosoma atrum Cast. Det. M.Baehr 2011” (ANIC). 16.4 mm
Copyright © Martin Baehr

Helluobrochus brevicollis (Dejean, 1831)
Male adult
Ecuador, Yasuni
Photo I.Sokolov
Copyright © Igor Sokolov

Helluomorpha eulinae Reichardt, 1974
Male adult
Paraguay, Dep. Pte. Hayes, Puerto Galileo,S25.04 W57.52 5-8.III.2008 leg. U.Drechsel. 19 mm; det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Macrocheilus saulcyi Chevrolat, 1854
Male adult
Libanon, Djezin, det. Ing. Jedlicka
Copyright © Ron Felix

Macrocheilus vanharteni Felix & Muilwijk, 2007 Holotype
Male adult
Seyun, 12-14.VIII.2002, light trap, leg. A.v. Harten &
A. Al Zubayri, Holotype, male 12,4 mm
Photo R. Felix
Copyright © Alexander Anichtchenko

Macrocheilus vanharteni Felix & Muilwijk, 2007 Paratype
Male adult
Al Kowd, VI.2002, light trap, leg. A.v.Harten & S. Al
Haruri, paratype, male, collection R. Felix
Photo R. Felix
Copyright © Alexander Anichtchenko

Helluomorphoides praeustus bicolor (T.W.Harris, 1828)
Male adult
Copyright Henri Goulet, Agriculture & Agri-Food Canada
Copyright © Henri Goulet

Macrocheilus bicolor Andrewes, 1920 Holotype
Female adult
Kaoeqaoe, 740, Type, Macrochilus bicolor Type Andr., H. E. Andrewes det.; H. E. Andrewes Coll. B. M. 1945–97. (NHML)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus impictus (Wiedemann, 1823)
Male adult
(MNHN)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus immanis Andrewes, 1920 Holotype
Male adult
Toungoo; Type; Macrochilus immanis, Type, Andr., H. E. Andrewes det. (NHML)<br /> photo Zhao & Tian, 2012<br />
Copyright © - - -

Macrocheilus niger Andrewes, 1920 Holotype
Female adult
type, 11, H. L. Andrewes, Nilgiri Hills, H. E. Andrewes Coll., B.M. 1945–97.; Macrochilus niger Type, Andr.,<br /> H. E. Andrewes det.; Macrochilus niger Andrewes, 1920 (deposited in NHML)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus asteriscus (White, 1844)
Male adult
photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus vitalisi Aandrewes, 1920 Holotype
Female adult
China, Bowring 63·47, 986, 27/2/53 (NHML)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus binotatus Andrewes, 1931
Female adult
Paggar Alam, Sumatra, J. Bouchaud (deposited in MNHN)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus gigas Zhao & Tian, 2010 Holotype
Male adult
Guangdong: Zhanjiang, Jun.1983 (deposited in SCAU)<br /> Photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus parvimaculatus Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Liuzhou: Luzhai, 26 May 1980, Shaozhou Ruan leg. (SCAU)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus tripustulatus (Dejean, 1825) Holotype
Female adult
tripustulatus Dejean (non Fabr.), Java, Coll. Dejean, 3-pustulatus Wiedemann (Helluo). Fabr. (Brachinus), Western, Ex-Musaeo Chaudoir (MNHN)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus chaudoiri Andrewes, 1919 Holotype
Female adult
Ex-Musaeo Chaudoir; Macrocheilus Chaudoiri Andr., H.S. Andrewes det.; 3-maculatus Chaud., Deccan, Coll. Jeakes (MNHN)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus nigrotibialis Heller, 1900 Holotype
Male adult
Drs. Sarasin N. Celebes Panot-Maimang, “12623”, “Staatl. Museum fur Tierkunde. Dresden” (deposited in SNSD)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus bensoni Hope, 1838
Male adult
photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus deuvie Zhao & Tian, 2012 Holotype
Male adult
Philippines, Bohol. Ch. Semper (deposited in MNHN)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus fuscipennis Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Napo, 10 Oct.1970, by light trap<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus solidipalpis Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Dibei, Oct.1980, Xiuzhen Mao leg., by light trap.<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus cheni Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Tengxian, Oct.1980<br /> Photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus quadratus Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Cangwu, 1980, in paddy field (SCAU)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus sinuatilabris Zhao & Tian, 2010 Holotype
Male adult
Guangxi: Fenghuangcheng, Sep.1981, by light trap (SCAU)<br /> photo Zhao & Tian, 2012
Copyright © - - -

Macrocheilus biguttatus Gory, 1832
Female adult
Senegal, M’Bour Nianing, piège lumineux 24/09/2009, Pascal Deschamps coll.
Copyright © Pascal Deschamps

Omphra atrata (Klug, 1834)
Male adult
India, Goa, Cotigao Wildlife Senctuary, 25.03.2013, P. & L. Ustjuzhanin (ISEA)
R. Dudko det. et photo, 2013
Copyright © Roman Dudko

Erephognathus ranomafanae Kavanaugh and Rainio, 2016 Holotype
Female adult
MADAGASCAR: Province Fianarantsoa, Parc National Ranomafana, Vohiparara, at broken bridge, el 1110 m 8-15 November 2001”/ “21°13.57’S, 047°22.19’E collector: R. Harin’Hala California Acad of Sciences malaise trap in high altitude rainforest, MA-02-09A-02”; HOLOTYPE Erephognathus ranomafanae Kavanaugh & Rainio sp. n. 2015 (CAS). Image from Kavanaugh & Rainio, 2016
Copyright © - - -

Colfax stewensi Andrewes, 1920
Male adult
10-11 mm Nekka Nalla, Ratnagiri District, Maharashtra, India, February 1973, M. Baburao leg. det. Danyang Zhao
Copyright © Shripad V. Manthen

Macrocheilus niger Andrewes, 1920
Male adult
16-16.50 mm Latitude 17.73625, Longitude 73.76983, Elevation (m) 671 Mhavashi, Satara District, Maharashtra, India, March 2017, leg. S.R Patil & Party; det. Shripad Manthen
Copyright © Shripad V. Manthen

Omphra complanata Reiche, 1843
Male adult
14 mm; GPS: Latitude 17.71172, Longitude 73.73987, Elevation (m) 773 Vengale, Satara District, Maharashtra, India, March 2017, leg. S.R Patil & Party; det. Shripad Manthen
Copyright © Shripad V. Manthen

Pleuracanthus sulcipennis Gray, 1832
Male adult
Bolivia, Dep. Santa Cruz, San Ignacio, 11.1996, leg. A. Mater (Collection R. Sciaky). 15 mm
Copyright © Alexander Anichtchenko

Meladroma angustata Basilewsky, 1970
Male adult
Rep. Centr. Africa, Maboke, M. Gombe, Doko, XI. 1985. (Collection and det by R. Sciaky as informicollis Liebke, 1928). 23mm ; det. Peter Schüle 2022
Copyright © Alexander Anichtchenko

Dailodontus clandestinus (Klug, 1834)
Male adult
Paraguay, Dep. Pte. Hayes, Gral. Bruguez, S24,44 W58,48 19-23.I.2006 leg. U. Drechsel. 12 mm. det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Helluomorpha eulinae Reichardt, 1974
Female adult
Paraguay, Dep Cordillera, Pirareta, 6-8.10.2012 U. Drechsel leg. 17 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Helluomorpha macroptera Chaudoir, 1850
Male adult
Paraguay, dep Guira, Ita Azul, 7-11.1.2011, U. Drechsel leg. det. A. Anichtchenko 2019
Copyright © Alexander Anichtchenko

Macrocheilus asteriscus (White, 1844)
Female adult
Laos
Copyright © Alexander Anichtchenko

Helluomorphoides squiresi (Chaudoir, 1872)
Male adult
Paraguay, Dep. Caazapa, Colonia Neufeld, 24.10-2.11.2008. 12 mm. det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Helluomorphoides rubricollis (Schaum, 1863)
Male adult
Paraguay, Dep. Caazapa, La Golondrina, S25,32 W55,29 14-19.III.2009 leg. U. Drechsel. det. & coll. Anichtchenko A.
Copyright © Alexander Anichtchenko

Helluobrochus darlingtoni Reichardt, 1974
Male adult
Paraguay, dep. Caazapa, Colonia Neufeld, S26.28 W55.55, 24.10.2008, leg. U. Drechsel 15 mm; det. Anichtchenko A. 2019
Copyright © Alexander Anichtchenko

Macrocheilus chinnarensis Akhil, Divya & Sabu, 2019 Holotype
Male adult
Holotype, Male, India: Kerala: Chinnar, 420 m, 08.viii.2016; coll. S.V. Akhil . Image from Akhil, Divya & Sabu, 2019
Copyright © - - -

Macrocheilus bandipurensis Akhil, Divya & Sabu, 2019 Holotype
Male adult
Holotype, Male, India: Karnataka: Gundlupet, 06.v.2016; coll. S.V. Akhil. Image from Akhil et al, 2019
Copyright © - - -

Omphra hirta (Fabricius, 1801)
Male adult
S India, Kerala, Trivandrum Dt., Poonmudi mg., 3000 ft, 5.1989 (cAA)
Copyright © Alexander Anichtchenko

Meladroma umbraculata (Fabricius, 1801)
Male adult
Cote d'Ivoire, Lamto Toumodi, 23-08-1971
Copyright © Pascal Deschamps

Omphra erwini Akhil & Sabu, 2021 Holotype
Male adult
India: Gujarat: Ambaji, 19.ii.1992, coll. ZSI-DRC, det. Akhil S. V. Image from Akhil & Sabu, 2021. Aedeagus deformed.
Copyright © - - -

Omphra balli Akhil & Sabu, 2021 Holotype
Male adult
India: Telangana: Amrabad TR: View point Farhabad, 19.ix.2018, coll. S. S. Jadhav, det. Akhil S. V. Image from Akhil & Sabu, 2021
Copyright © - - -

Macrocheilus saulcyi Chevrolat, 1854
Female adult
Jordan, Petra (cAA). det. A. Anichtchenko, 2021
Copyright © Alexander Anichtchenko

Macrocheilus cambodianus Allegro & Giachino, 2021 Holotype
Male adult
Cambodia, Banteay Meanchey Province, near Sisophon, campus of the Mean Chey University, 13°14’45”n, 20°56’29”E, light trap, 22.x-23.xI.2019, P. Bun & V. Kong leg. (CAl).
Copyright © Gianni Allegro

Macrocheilus walterrossii Allegro & Giachino, 2021 Holotype
Male adult
Cambodia, Banteay Meanchey Province, near Sisophon, campus of the Mean Chey University, 13°14’45”n, 20°56’29”E, light trap, 22.x-23.xI.2019, P. Bun & V. Kong leg. (CGi).
Copyright © Gianni Allegro

Triaenogenius (Triaenogenius) lugubrinus (Boheman, 1860)
Male adult
SW Africa, Kaokoveld, Sanitatas abr 85 miles WSW Ohopoho 14-16.6.1951 Swedish Sout Africa Expedition (cAA) 18 mm
Copyright © Alexander Anichtchenko

Macrocheilus biguttatus Gory, 1832
Female adult
Afrique de L`Ouest, Senegal Petit Cote, Mbour 20.IX.2009 IRD, Ultra Violet, Alain Coache
Copyright © Pascal Deschamps

Macrocheilus biguttatus Gory, 1832
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus allardi Basilewsky, 1957 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus alluaudi Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus alluaudi Burgeon, 1937
Male adult
Republic of Congo, Réserve Naturelle des Gorilles de Lésio Louna 17–28.11.2017, Batéké Plateau H. Krahnstôver leg. (Collection P. Schüle, Herrenberg). Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus assmanni Serrano, 2023 Holotype
Female adult
Tanzania, 12-17.XII.1996\ Ruvuma, near Songea\ leg. Werner & Lizler. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus angustatus Basilewsky, 1949 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus basilewskyi A.Serrano, 2000 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus bimaculatus (Dejean, 1831)
Male adult
Quebo (Coli), 28/06/1995, Col. A. Serrano\\ Miss. Ent. SPEN, à Guiné-Bissau, 25 Junho a 17 Julho 95. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus biplagiatus (Boheman, 1848)
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus burgeoni Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus clasispilus Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus crampeli Alluaud, 1916 Paratype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus cruciatus (Marc, 1840)
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus diplospilus Basilewsky, 1967 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus dorsalis Klug, 1834
Male adult
Zambia NC., 82 km SSW Solwezi, 3.12.2004, Snižek, Tichẏ lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus dorsiger (Chaudoir, 1876)
Male adult
Burkina Faso: Nahouri, Forêt de Nazinga, Akwazena, 275 m, 11º 09′ 24″ N 001º 36′ 44″ O, 21.VII.2006, zone Soudanienne, savane boisée, piège lumineux, F & S. Génien, 2006-53. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus elegantulus Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus hybridus Peringuey, 1896
Female adult
Zimbabwe W Bulawayo, Shangani, Naletale ruins env. 20.12.2011, Snižek. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus kudrnai Serrano, 2023 Holotype
Male adult
NE Benin\ E of Kandi\ 4 km W of Bensekou\ 24–25.06.2001\ A. Kudrna Jr. Lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus labrosus (Dejean, 1831)
Male adult
Central African Republic. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus longicollis Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus longicollis Peringuey, 1904
Male adult
Zambia NE 2004, 30–60 km NW of Mpika 24.11. Snižek, Tichẏ lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus longicollis Peringuey, 1904
Male adult
Moçambique, 14.XII.2003 near Dombe, Werner & Lizler leg. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus moraisi A.Serrano, 2000 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus ocellatus Basilewsky, 1953 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus overlaeti Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus persimilis Basilewsky, 1970 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus persimilis Basilewsky, 1970
Male adult
Zambia, Northwestern Prov. 20 km SE Mwinilunga 7–8.01.2003 A. Kudrna Jr. Lgt. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus philippeorum Serrano, 2023 Holotype
Male adult
Burkina Faso, Pama\ 24–28.VIII.2005,\ 11º 17′ 00′′ N, 0º 42′59E, 232m\ Phillippe Moretto leg. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus proximus Peringuey, 1896 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus proximus Peringuey, 1896
Female adult
Ethiopia, 25/27.IV.1998\ near Negele Borana\ Sidamo, leg. Werner. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus quadrinotatus Burgeon, 1937 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus quadrinotatus Burgeon, 1937
Female adult
Kibundi (15 km N Nambua). Malanje, Angola, 19.XI.2017, 8º 54′ 33.31′′ S 16º 3′ 15.71′′ E, 1170 m. A. Serrano & R. Capela leg. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus scapularis Reiche, 1843
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus schuelei Serrano, 2023 Holotype
Male adult
RCA, P.N. Ndoki, Camp 1, Chablis 1,\ 02º 28′ 40.5N 016º 13′ 02.6E, 24–25.II.\ 2012 forêt diversifiée, piège UV. Exp.\ Sangha 12, P. Moretto leg. -117-“ \\ “Collection P. Schüle, Herrenberg\ Shanga Projekt 2012” . Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus spectandus Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus spectandus Peringuey, 1904
Male adult
Tanzania, 9/13.XII.1997 near Songea, Ruvuma Prov. leg. Werner & Lizler. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus vinctus Basilewsky, 1960 Paratype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus taedatus Basilewsky, 1960 Paratype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus taedatus Basilewsky, 1960
Female adult
Tanzania, 10/16.XII.99, near Mitundo, Dodoma prov. Werner & Lizler leg. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus viduatus katanganus Basilewsky, 1960 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus viduatus katanganus Basilewsky, 1960
Female adult
Zambia. Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus varians eugrammus Basilewsky, 1960 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Macrocheilus varians Peringuey, 1904 Holotype
Male adult
Image from A. Serrano, 2023
Copyright © - - -

Helluomorphoides texanus (Leconte, 1853)
Male adult
U.S.A. Texas Comté de Parker Dennis 29-06-1985
Copyright © Pascal Deschamps

Colfax stevensi Andrewes, 1920
Male adult

Copyright © Jong-Bong Choi

Colfax stevensi Andrewes, 1920
Male adult

Copyright © Jong-Bong Choi

Macrocheilus varians eugrammus Basilewsky, 1960
Male adult
RSA, Limpopo prov., SW of Sekgopo, Mahulo lodge, 23*658S 29*881E 12.12.2023 Bednarik leg. (collection Rudolf Kmeco). det. Anichtchenko A. 2025. 10 mm
Copyright © Alexander Anichtchenko