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Carabidae of the World
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Carabinae Latreille, 1802
→
Carabini Latreille, 1802
→
Carabina sensu stricto
→
Carabus Linne, 1758
→
Procerus Dejean, 1828
→ scabrosus Olivier, 1795
Region
Afghanistan (AF)
Albania (AL)
Aleutian isl (ALU)
Algeria (AG)
Andorra (AN)
Angola (ANG)
Antarctica (Antarctica)
Antilles islands (ANT)
Antigua and Barbuda (Antigua and Barbuda)
Barbados (Barbados)
Cayman islands (Cayman islands)
Cuba (Cuba)
Dominica (Lesser Antilles) (Dominica (Lesser Antilles))
Dominican Republic (Hispaniola, Greater Antilles) (Dominican Republic (Greater Antilles))
Grenada (Grenada)
Guadeloupe (Guadeloupe)
Haiti (Haiti)
Jamaica (Jamaica)
Martinique (Martinique)
Puerto Rico (Puerto Rico)
Saint Kitts and Nevis (Saint Kitts and Nevis)
Saint Lucia (Saint Lucia)
St. Vincent (St. Vincent)
Trinidad and Tobago (Trinidad and Tobago)
Arab Emirates (AE)
Argentina (ARG)
Armenia (AR)
Australia (AST)
Christmas Island (Christmas Island)
Lord Howe Island (Lord Howe Island)
Macquarie Island (Macquarie Island)
Tasmania (Tasmania)
Austria (AU)
Azerbaijan (AB)
Azores (AZ)
BALTIC AMBER, Baltic See coast and Amber quarry Jantarny near Kaliningrad, Russia (BAM)
Bahamas (BHM)
Bahrain (BA)
Bangladesh (BGD)
Belgium (BE)
Belize (BLZ)
Bhutan (BT)
Bolivia (BOL)
Bosnia and Herzegovina (BH)
Botswana (BTW)
Brazil (BRZ)
Acre (AC)
Alagoas (AL)
Amazonas (AM)
Arnapá (AP)
Bahia (BA)
Ceará (CE)
District Federal (Brasilia) (DF)
Espirito Santo (ES)
Goiás (GO)
Maranhão (MA)
Mato Grosso (MT)
Mato Grosso do Sul (MS)
Minas Gerais (MG)
Paraná (PR)
Paraîba (PB)
Pará (PA)
Pernambouc (PE)
Piaui (PI)
Rio Grande do Norte (RN)
Rio Grande do Sul (RS)
Rio de Janeiro (RJ)
Rondônia (RO)
Roraima (RR)
Santa Catarina (SC)
Sergipe (SE)
São Paulo (SP)
Tocantins (TO)
Brunei (Brunei)
Bulgaria (BU )
Burundi (BUR)
Byelorussia (BY)
Cambodia (CBD)
Cameroon (CAM)
Canada (CAN)
Canary isls (CI)
Cape Verde (Cape Verde)
Central African Republic (CAR)
Chad (CHD)
Chile (CHL)
Juan Fernández Islands (Juan Fernández Islands)
China (CH)
Anhui (Anhui)
Beijing (Beijing)
Chongqing (Chongqing)
Fujian (Fujian)
Gansu (Gansu)
Guangdong (Guangdong)
Guangxi (Guangxi)
Guizhou (Guizhou)
Hainan (Hainan)
Hebei (Hebei)
Heilongjiang (Heilongjiang)
Henan (Henan)
Honkong (Hongkong)
Hubei (Hubei)
Hunan (Hunan)
Inner Mongolia (Inner Mongolia)
Jiangsu (Jiangsu)
Jiangxi (Jiangxi)
Jilin (Jilin)
Liaoning (Liaoning)
Macau (Macau)
Ningxia (Ningxia)
Qinghai (Qinghai)
Shaanxi (Shaanxi)
Shandong (Shandong)
Shanghai (Shanghai)
Shanxi (Shanxi)
Sichuan (Sichuan)
Tianjin (Tsiensin) (Tianjin)
Xinjiang (Xinjiang)
Xizang (Tibet) (Xizang (Tibet))
Yunnan (Yunnan)
Zhejiang (Zhejiang)
Colombia (COL)
Comoros isl. (COM)
Costa Rica (CRC)
Croatia (CR)
Crozet Islands (CRZ)
Cyprus (CY)
Czech (CZ)
DOMINICAN AMBER, Dominican Republic, Lower Miocene. (DAM)
Democratic Republic of the Congo (CNG)
Denmark (DE)
Diego Garcia isls (DGC)
Djibouti (DJB)
Ecuador (ECD)
Egypt (EG)
Sinai (Sinai)
El Salvador (SVD)
Equatorial Guinea (EQG)
Bioco (Fernando Pó) island (Bioco (Fernando Pó) island)
Eritrea (ERT)
Estonia (EN)
Ethiopia (ETP)
Falkland il. (FLK)
Faroe Islands (FA)
Finland (FI)
France (FR)
Corse (Corse)
French Guiana (GNF)
Gabon (GAB)
Galapagos Islands (GAL)
Georgia (GG)
Abkhazia (Abkhazia)
Ajaria (Ajaria)
South Ossetia (South Ossetia)
Germany (GE)
Great Britain (GB)
Greece (GR)
Aegean Islands (Aegean Islands)
Crete (Crete)
Dodecanese Islands (Dodecanese Islands)
Euboea (Evia) Island (Euboea (Evia) Island)
Ionian Islands (Ionian Islands)
Peloponnese (Peloponnese)
Saronic Islands (Saronic Islands)
Sporades Islands (Sporades Islands)
The Cyclades (The Cyclades)
Guatemala (GTM)
Guyana (GYN)
Honduras (HDR)
Hungary (HU)
Iceland (IC)
India (IND)
Andaman and Nicobar isl. (Andaman and Nicobar isl.)
Andhra Pradesh (Andhra Pradesh)
Arunachal Pradesh (Arunachal Pradesh)
Assam (Assam)
Bihar (Bihar)
Chhattisgarh (Chhattisgarh)
Delhi (Delhi)
Goa (Goa)
Gujarat (Gujarat)
Haryana (Haryana)
Himachal Pradesh (Himachal Pradesh)
Jammu and Kashmir (Jammu and Kashmir)
Jharkhand (Jharkhand)
Karnataka (Karnataka)
Kerala (Kerala)
Ladakh (Ladakh)
Madhya Pradesh (Madhya Pradesh)
Maharashtra (Maharashtra)
Manipur (Manipur)
Meghalaya (Meghalaya)
Mizoram (Mizoram)
Nagaland (Nagaland)
Odisha (Odisha)
Puducherry (Puducherry)
Punjab (Punjab)
Rajasthan (Rajasthan)
Sikkim (Sikkim)
Tamil Nadu (Tamil Nadu)
Telangana (Telangana)
Tripura (Tripura)
Uttar Pradesh (Uttar Pradesh)
Uttarakhand (Uttarakhand)
West Bengal (West Bengal)
Indonesia (IDS)
Borneo (Kalimantan) (Borneo)
Java (Java)
Lesser Sunda Islands (Lesser Sunda Ils.)
Maluku Islands (Maluku Isl.)
Papua (Papua)
Riau Islands (Riau Islands)
Sulawesi (Celebes) (Sulawesi)
Sumatra (Sumatra)
Timor (Timor)
Iran (IN)
Iraq (IQ)
Ireland (IR)
Israel (IS)
Italy (IT)
Sardegna (Sardegna)
Sicily (Sicily)
Japan (JA)
Hokkaido (Hokkaido)
Honshu (Honshu)
Kyushu (Kyushu)
Ryukyu Islands (Ryukyu Islands)
Shikoku (Shikoku)
Jordan (JO)
Kashmir (KA)
Kazakhstan (KZ)
Kenya (KEN)
Kerguelen Islands (Kerguelen Islands)
Kuwait (KU)
Kyrgyzstan (KI)
Laos (LAO)
Latvia (LA)
Lebanon (LE)
Lesotho (LSO)
Libya (LB)
Liechtenstein (LS)
Lithuania (LT)
Luxenbourg (LU)
Macedonia (MC)
Madagascar (MAD)
Madeira Arch. (MR)
Malawi (MLW)
Malaysia (MLS)
Sabah (Borneo) (Sabah (Borneo))
Sarawak (Borneo) (Sarawak (Borneo))
Maldives (MLD)
Malta (MA)
Mauritania (MAU)
Mauritius (MRC)
Melanesia (Melanesia)
Bismarck Archipelago (Bismarck Archipelago)
Fiji (Fiji)
New Caledonia (New Caledonia)
Papua New Guinea (Papua New Guinea)
Solomon Islands (Solomon Islands)
Vanuatu ils (Vanuatu ils)
Mexico (MEX)
Aguascalientes (Aguascalientes)
Baja California (Baja California)
Baja California Sur (Baja California Sur)
Campeche (Campeche)
Chiapas (Chiapas)
Chihuahua (Chihuahua)
Coahuila (Coahuila)
Colima (Colima)
Durango (Durango)
Federal District (Federal District)
Guanajuato (Guanajuato)
Guerrero (Guerrero)
Hidalgo (Hidalgo)
Jalisco (Jalisco)
Mexico State (Mexico State)
Michoacan (Michoacan)
Morelos (Morelos)
Nayarit (Nayarit)
Nuevo Leon (Nuevo Leon)
Oaxaca (Oaxaca)
Puebla (Puebla)
Queretaro (Queretaro)
Quintana Roo (Quintana Roo)
San Luis Potosi (San Luis Potosi)
Sinaloa (Sinaloa)
Sonora (Sonora)
Tabasco (Tabasco)
Tamaulipas (Tamaulipas)
Tlaxcala (Tlaxcala)
Veracruz (Veracruz)
Yucatan (Yucatan)
Zacatecas (Zacatecas)
Micronesia (MIC)
Caroline islands (Caroline islands)
Guam (Guam)
Kiribati (Kiribati)
Marshall ils (Marshall ils)
North Marians isl. (North Marians isl.)
Palau archipelago (Palau archipelago)
Moldavia (MD)
Mongolia (MG)
Montenegro (ME)
Mordor (RU)
Altai Republic and Altaisky Province (Altai Republic and Altaisky Province)
Amurskaya Area (Amurskaya Area)
Bashkiria, E of Chelyabinsk regs. (S Ural mts. Bashkiria, E of Chelyabinsk regs.)
Buryat Republic (Buryat Republic)
Chukot autonomous region (Chukotka)
East Sayan Mountains (East Sayan Mountains)
Ingushetia, Chechnya, Daghestan (E Caucas) (Ingushetia, Chechnya, Daghestan (E Caucas))
Irkutsk Area (Irkutsk Area)
Kabardino-Balkaria, Ossetia (C Caucas) (Kabardino-Balkaria, Ossetia (C Caucas))
Kaliningrad, Briansk, Smolensk, Tula, Orlov, Kaluga, Moskow, Riaz (C of Russian plains)
Kamchatskaya Area (Kamchatka)
Karachaevo-Cherkessia (W Caucas) (Karachaevo-Cherkessia)
Karelia, Leningrad reg., Pskov reg., Kostroma reg., Perm reg., Komi, Pechora plains. (N of Russian plains)
Khabarovsky Province (Khabarovsky Province)
Khamar-Daban Mountains (Khamar-Daban Mountains (E Siberia))
Komi, Yamalo-Nenetsk, Hanty-Mansi (N Ural mts.)
Krasnodar reg. (W Caucas)
Krasnoyarsky Province (Krasnoyarsk)
Kuriles Islands (Kuriles Islands)
Magadanskaya Area (Magadan)
Omsk, S of Tyumen, W Novosibirsk regs. (S of W Siberia)
Perm, Sverdlovsk regs. (C Ural mts)
Primorsky District (Primorie) (Maritime Prov)
Sakhalin Island (Sakhalin Island)
Tuva Republic (Tuva)
Voronezh, Rostov, Saratov, Kalmykia, Volgograd, Astrahan regs. (S of Russian plains)
West Sayan Mountains (West Sayan)
Yakut-Saha Republic (Yakut-Saha Republic)
Zabaykalsky Krai (ex Chita reg.) (Zabaykalsky Krai)
plains of Krasnodar reg., S Rostov reg., Stavropol reg. (Ciscaucas reg. (plains of Krasnodar reg., S Rostov)
taiga from Ural mts to Yenisei riv. (Middle of W Siberia)
tundra of Tyumen, Yamalo-Nenetsk regs. (N of W Siberia)
Morocco (MO)
Mozambique (MZB)
Myanmar (Burma) (BRM)
Namibia (NAM)
Nepal (NP)
Netherlands (NL)
New Zealand (NZL)
Antipodes Islands (Antipodes Islands)
Auckland Islands (Auckland Islands)
Campbell Island (Campbell Island)
Chatham Islands (Chatham Islands)
Nicaragua (NCG)
North Korea (NC)
Norway (NR)
Svalbard (Spitzbergen) (SB)
Oman (OM)
Pakistan (PA)
Panama (PNM)
Paraguay (PRG)
Peru (PER)
Philippines (PLP)
Basilan (Basilan)
Cebu (Cebu)
Guimaras (Guimaras)
Leyte (Leyte)
Luzon (Luzon)
Marinduque (Marinduque)
Masbate (Masbate)
Mindanao (Mindanao)
Mindoro (Mindoro)
Negros (Negros)
Palawan (Palawan)
Panay (Panay)
Polillo Islands (Polillo Islands)
Romblon (Romblon)
Samar (Samar)
Sulu Archipelago (Sulu Archipelago)
Poland (PL)
Polynesia (POL)
Cook Islands (Cook Islands)
Hawaiian Islands (HAW)
Marquesas Islands (Marquesas Islands)
Niue (Niue)
Norfolk ils (Norfolk ils)
Pitcairn ils (Pitcairn ils)
Rapa Nui (Easter Island) (Rapa Nui)
Samoa (Samoa)
Society Islands (Society Islands)
Tahiti (Tahiti)
Tokelau (TKL)
Tonga (Tonga)
Tuvalu ils (Tuvalu ils)
Portugal (PT)
Qatar (QA)
Republic of South Africa (RSA)
Republic of the Congo (CG)
Reunion isl. (Reunion isl.)
Rodriguez isls. (ROD)
Romania (RO)
Rwanda (RWA)
Santa Helena isl (SH)
Sao Tome e Principe (STM)
Saudi Arabia (SA)
Serbia (SRB)
Seychelles (SEY)
Singapore (SGP)
Slovakia (SK)
Slovenia (SL)
Somalia (SOM)
South Georgia and the South Sandwich Islands (SGSSI)
South Korea (SC)
Spain (SP)
Balearic Islands (Balearic Islands)
Sri Lanka (SRL)
St. Paul isls (STP)
Sudan (SUD)
Surinam (SUR)
Swaziland (SWZ)
Sweden (SV)
Switzerland (SZ)
Syria (SY)
Tadzhikistan (TD)
Taiwan (Formosa) (Taiwan)
Tanzania (TAN)
Zanzibar isl. (Zanzibar isl.)
Thailand (TAI)
Tunisia (TU)
Turkey (TR)
Turkmenistan (TM)
Uganda (UGD)
Ukraine (UK)
Crimea (Crimea)
United States (USA)
Alabama (Alabama)
Alaska (Alaska)
Arizona (Arizona)
Arkansas (Arkansas)
California (California)
Colorado (Colorado)
Connecticut (Connecticut)
Delaware (Delaware)
Florida (Florida)
Georgia (Georgia)
Idaho (Idaho)
Illinois (Illinois)
Indiana (Indiana)
Iowa (Iowa)
Kansas (Kansas)
Kentucky (Kentucky)
Louisiana (Louisiana)
Maine (Maine)
Maryland (Maryland)
Massachusetts (Massachusetts)
Michigan (Michigan)
Minnesota (Minnesota)
Mississippi (Mississippi)
Missouri (Missouri)
Montana (Montana)
Nebraska (Nebraska)
Nevada (Nevada)
New Hampshire (New Hampshire)
New Jersey (New Jersey)
New Mexico (New Mexico)
New York (New York)
North Carolina (North Carolina)
North Dakota (North Dakota)
Ohio (Ohio)
Oklahoma (Oklahoma)
Oregon (Oregon)
Pennsylvania (Pennsylvania)
Rhode Island (Rhode Island)
South Carolina (South Carolina)
South Dakota (South Dakota )
Tennessee (Tennessee)
Texas (Texas)
Utah (Utah)
Vermont (Vermont)
Virginia (Virginia)
Washington (Washington)
West Virginia (West Virginia)
Wisconsin (Wisconsin)
Wyoming (Wyoming)
Uruguay (URG)
Uzbekistan (UZ)
Venezuela (VNZ)
Vietnam (VTN)
West Africa (WAF)
Benin (Benin)
Burkina Faso (Burkina Faso)
Côte d'Ivoire (Côte d'Ivoire)
Gambia (Gambia)
Ghana (Ghana)
Guinea (Guinea)
Guinea Bissau (Guinea Bissau)
Liberia (Liberia)
Mali (Mali)
Niger (Niger)
Nigeria (Nigeria)
Senegal (Senegal)
Sierra Leone (Sierra Leone)
Togo (Togo)
Yemen (YE)
Socotra (Socotra)
Zambia (ZMB)
Zimbabwe (ZIM)
Main
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Adult
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Labels
Carabus (Procerus) scabrosus akbesianus Breuning, 1975
Male adult
Akbés (Antakya, Turquey)
Copyright ©
Alain Mollard
Carabus (Procerus) scabrosus akbesianus Breuning, 1975
Holotype
Male adult
Syrie, Akbes (MNHN-EC-EC131)
Copyright © - - -
akbesianus
taxon page
(2 photos)
Carabus (Procerus) scabrosus amasicus Csiki, 1927
Male adult
Almus (Tokat, Turquey)
Copyright ©
Alain Mollard
amasicus
taxon page
(1 photos)
Carabus (Procerus) scabrosus armenius Zaitzev, 1930
Male adult
Balkaya (Gumushane, Turquey)
Copyright ©
Alain Mollard
armenius
taxon page
(1 photos)
Carabus (Procerus) scabrosus audouini Brulle, 1835
Male adult
Isiklar (Trabzon,Turquey)
Copyright ©
Alain Mollard
Carabus (Procerus) scabrosus audouini Brulle, 1835
Female adult
Turkey. Rize Prov., South of Ikizdere, 800 m, 17.08.2009. Image of В.И. Гуменюк from www.zin.ru
Copyright © - - -
audouini
taxon page
(2 photos)
Carabus (Procerus) scabrosus bureschianus Breuning, 1928
Male adult
Bulgaria, W Rodopi, Dobrostan masive, backovo env. M.Janalik leg.
Copyright ©
Igor Solodovnikov
bureschianus
taxon page
(1 photos)
Carabus (Procerus) scabrosus callipygius Cavazzuti, 1986
Male adult
Devrekani (Kastamonu, Turquey)
Copyright ©
Alain Mollard
callipygius
taxon page
(1 photos)
Carabus (Procerus) scabrosus caucasicus Adams, 1817
Male adult
N Caucasus,
Stavropol Krai, Pyatigorsk, Mt.Mashuk, VI.2007 Tikhonov Yu. leg.
44mm
Copyright ©
Ruslan Panin
Carabus (Procerus) caucasicus var.tuberculosus
Lectotype
Female adult
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC468 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec468
Copyright © - - -
Carabus (Procerus) scabrosus var.tuberculosus Géhin, 1885
Label
Muséum national d’Histoire naturelle, Paris (France) Collection : Insects: Coleoptera (EC) Specimen MNHN-EC- EC468 http://coldb.mnhn.fr/catalognumber/mnhn/ec/ec468
Copyright © - - -
caucasicus
taxon page
(3 photos)
Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
Male adult
Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov
Copyright ©
Igor Solodovnikov
Carabus (Procerus) scabrosus colchicus Motschulsky, 1844
Male adult
Abkhasia: S part of Bzybian Mt.R. vall. riv. Khipsta, h= 600 m, VIII.2014, leg. I.A. Solodovnikov, 58 mm
Copyright ©
Igor Solodovnikov
Carabus (Procerus) scabrosus tatianagorokhovae Cavazzuti & Kozlov, 2014
Holotype
Male adult
Armenia. Lori prov. Akhtala
Copyright ©
Anton Kozlov
Carabus (Procerus) scabrosus vardziaensis Deuve, 2017
Holotype
Female adult
Copyright ©
Ruslan Panin
colchicus
taxon page
(4 photos)
Carabus (Procerus) scabrosus dardanellicus Kraatz, 1886
Male adult
Sancaeli (Canakkale, Turquey)
Copyright ©
Alain Mollard
dardanellicus
taxon page
(1 photos)
Carabus (Procerus) scabrosus estegeicus (Cavazzuti, 1989)
Female adult
Copyright ©
Özgür KOÇAK
estegeicus
taxon page
(1 photos)
Carabus (Procerus) scabrosus fallettianus (Cavazzuti, 1997)
Male adult
Khndzorek (Goris, Arménie)
Copyright ©
Alain Mollard
Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014
Paratype
Female adult
Tavush prov., Ditavan vill. 25-29.VII.2013 (in coll Kozlov)
Copyright ©
Anton Kozlov
Carabus (Procerus) scabrosus antonkozlovi Cavazzuti, 2014
Holotype
Male adult
Tavush prov., Mehrab vill. 950-1005m (in coll Kozlov). 44 mm
Copyright ©
Anton Kozlov
Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016
Paratype
Female adult
Copyright ©
Anton Kozlov
Carabus (Procerus) scabrosus nakagomei Cavazzuti & Kozlov, 2016
Holotype
Male adult
Copyright ©
Anton Kozlov
fallettianus
taxon page
(5 photos)
Carabus (Procerus) scabrosus mentor Blumenthal & Breuning, 1967
Male adult
Catalzeytin,(Katsamonu,Turquey)
Copyright ©
Alain Mollard
mentor
taxon page
(1 photos)
Carabus (Procerus) scabrosus montisabanti Schweiger, 1962
Male adult
Abant (Bolu, Turquey)
Copyright ©
Alain Mollard
montisabanti
taxon page
(1 photos)
Carabus (Procerus) scabrosus munzurensis Cavazzuti & Lassalle, 1987
Paratype
Male adult
leg.B.Lassalle, from Pulumur (Tunceli,Turquey)
Copyright ©
Alain Mollard
munzurensis
taxon page
(1 photos)
Carabus (Procerus) scabrosus propinquus Csiki, 1927
Male adult
Alem Dag (Istambul, Turquey) L=57 mm
Copyright ©
Alain Mollard
propinquus
taxon page
(1 photos)
Carabus (Procerus) scabrosus scabrosus Olivier, 1795
Male adult
Dermirkoy (Kirklareli, Turquey). L = 54mm
Copyright ©
Alain Mollard
scabrosus
taxon page
(1 photos)
Carabus (Procerus) scabrosus schuberti Breuning, 1968
Male adult
NC-Turkey: 10 km E of Sinop, h = 0 m, VI.1999, leg. St. Remond, 42 mm
Copyright ©
Igor Solodovnikov
schuberti
taxon page
(1 photos)
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Female adult
Copyright ©
Safronov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
f. virens Crimea, Kazu-Kau Mt. (South part Chatyr-Dagh Mt.) 14.06.2009. Leg V.V.Savchuk, male. 41 mm
Copyright ©
Igor Solodovnikov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
f. purpureus Kraatz Crimea, Inkerman, steppe, 26.03.1987. Leg I.M. Lutzky. 42 mm
Copyright ©
Igor Solodovnikov
Carabus (Procerus) scabrosus tauricus Bonelli, 1811
Male adult
Crimea, Sevastopol, leg.det. A.Chuvilin, 12.VIII.1999. Photo: U.Schmidt, 2016. 40mm
Copyright ©
Udo Schmidt
tauricus
taxon page
(4 photos)
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